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Invertebrates behaviour winner: The Architectural Army by Daniel Kronauer, US. At dusk, Kronauer tracked the colony of nomadic army ants as it moved, travelling up to a quarter of a mile through the rainforest near La Selva biological station, in north-eastern Costa Rica. The ants would use their bodies to build a new daytime nest to house the queen and larvae. They would form a scaffold of vertical chains by interlocking claws on their feet and then create a network of chambers and tunnels into which the larvae and queen would be moved from the last bivouac. (Photo by Daniel Kronauer/2019 Wildlife Photographer of the Year)
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