Winner. “I took this vertical image in the Quarry Bay district of Hong Kong during the dusk ‘blue hour’, when there was a perfect balance between the ambient light in the sky and the artificial lights of the high-rise residential buildings”. (Photo by Jatinder Heer/The Guardian)

Behaviour winner; Courting Royals: two royal terns in courtship display by Kristian Bell. Another beautiful morning on a beautiful beach on the Gulf Coast of Florida seemed to prompt these two royal terns to commence an intricate courtship dance. The photograph was taken with a Canon 300mm lens and 2x extender. (Photo by Kristian Bell/Deakin University/Royal Society Publishing Photography Competition 2018)

Runner-up. “The heavens opened and the streets around Shibuya, Tokyo, were suddenly drenched in even more colour and movement. I was taking photos (my friend was holding the umbrella) when I noticed this woman at the crossing. I’m pleased with the futuristic feel of the image”. (Photo by Katherine Bridgestock/The Guardian)

A gravedigger competes as a boy looks on during the first National Grave Digging competition at the public cemetery of Debrecen, 226 km east of Budapest, Hungary, Friday, June 3, 2016. Eighteen two-man teams of Hungarian gravediggers are demonstrating their skills for a place in a regional championship to be held in Slovakia. (Photo by Zsolt Czegledi/MTI via AP Photo)

A swimmer reacts after finishing his competition in a pool carved into thick ice covering the Songhua River during the Harbin Ice Swimming Competition in the northern city of Harbin, Heilongjiang province January 5, 2015. The swimming competition was held on the official launch day of the Harbin International Ice and Snow Sculpture Festival. (Photo by Kim Kyung-Hoon/Reuters)

A greater bird of paradise (Paradisaea apoda) displaying in Badigaki Forest, Wokam Island (Aru Islands, Indonesia). Found here in Aru and on adjacent New Guinea, the greater bird of paradise represents about 40 different species of birds of paradise that depend on intact rainforest across the New Guinea region spanning eastern Indonesia and Papua New Guinea. With more than 80% of forest cover still intact, this region represents the largest remaining block of rainforest in the entire Asia-Pacific. (Photo by Tim Laman/naturepl.com/LDY Agency)

Mariusz Zbigniew Pudzianowski of Poland leads a match of the 2005 World's Strongest Man Competition at Wuhou Temple on September 27, 2005 in Chengdu of Sichuan Province, southwest China. (Photo by China Photos/Getty Images)

About a dozen dogs traded in their leashes for life vests Saturday for the chance to ride the waves at Jupiter beach as part of a canine surfing competition to raise money for an animal rescue group. Photo: Musso paddles in with Jengo. (Photo by Taylor Jones/The Palm Beach Post)