Winner. “I took this vertical image in the Quarry Bay district of Hong Kong during the dusk ‘blue hour’, when there was a perfect balance between the ambient light in the sky and the artificial lights of the high-rise residential buildings”. (Photo by Jatinder Heer/The Guardian)

Behaviour winner; Courting Royals: two royal terns in courtship display by Kristian Bell. Another beautiful morning on a beautiful beach on the Gulf Coast of Florida seemed to prompt these two royal terns to commence an intricate courtship dance. The photograph was taken with a Canon 300mm lens and 2x extender. (Photo by Kristian Bell/Deakin University/Royal Society Publishing Photography Competition 2018)

Andrey Karr from Western Riders Slacklines at sunset above big waves in Nazare, Portugal on December 27, 2017. (Photo by Aidan Williams/National Geographic Travel Photographer of the Year Contest)

A model presents a creation during the 2018 Victoria's Secret Fashion Show in New York City, New York, U.S., November 8, 2018. (Photo by Mike Segar/Reuters)

Runner-up. “The heavens opened and the streets around Shibuya, Tokyo, were suddenly drenched in even more colour and movement. I was taking photos (my friend was holding the umbrella) when I noticed this woman at the crossing. I’m pleased with the futuristic feel of the image”. (Photo by Katherine Bridgestock/The Guardian)

Max Niedermaier of Germany collides with uncontrolled motorcycle of Niclas Svensson of Sweden during Astana Expo FIM Ice Speedway Gladiators World Championship at the Medeo rink in Almaty, Kazakhstan, February 18, 2017. (Photo by Shamil Zhumatov/Reuters)

Riot police runs as a woman falls down during a May Day rally in the Kreuzberg district of Berlin, Germany, 01 May 2016. (Photo by Kay Nietfeld/EPA)

Spectators pass through security screening ahead of the New Year's Eve celebration in Times Square in New York, on Sunday, December 31, 2017. (Photo by Peter Morgan/AP Photo)