Behaviour winner; Courting Royals: two royal terns in courtship display by Kristian Bell. Another beautiful morning on a beautiful beach on the Gulf Coast of Florida seemed to prompt these two royal terns to commence an intricate courtship dance. The photograph was taken with a Canon 300mm lens and 2x extender. (Photo by Kristian Bell/Deakin University/Royal Society Publishing Photography Competition 2018)

Honourable mention, Behaviour category. Toss the scorpion – Indian roller playing with its kill by Susmita Datta. The image was taken during an early morning safari drive at Tadoba Andhari Tiger Reserve in India. (Photo by Susmita Datta/PA Wire/Royal Society Publishing Photography Competition 2017)

Runner-up. “The heavens opened and the streets around Shibuya, Tokyo, were suddenly drenched in even more colour and movement. I was taking photos (my friend was holding the umbrella) when I noticed this woman at the crossing. I’m pleased with the futuristic feel of the image”. (Photo by Katherine Bridgestock/The Guardian)

A Syrian refugee man (obscured) moves boxes of goods at his shop in Zaatari refugee camp near the border with Syria, in Mafraq, Jordan October 15, 2016. (Photo by Ammar Awad/Reuters)

A Palestinian man walks down snow-covered stairs after Friday prayers near the Dome of the Rock on the compound known to Muslims as Noble Sanctuary and to Jews as Temple Mount, in Jerusalem's Old City February 20, 2015. Snow covered Jerusalem and mountainous areas of Israel early Friday morning and the education ministry closed schools for the day. (Photo by Ammar Awad/Reuters)

An Israeli policeman prevents a Palestinian man from entering the compound which houses al-Aqsa mosque, known by Muslims as the Noble Sanctuary and by Jews as the Temple Mount, in Jerusalem's Old City September 28, 2015. (Photo by Ammar Awad/Reuters)

Police detain a woman during a protest against Prime Minister Benjamin Netanyahu and his government's handling of the coronavirus disease (COVID-19) crisis, near Netanyahu's residence in Jerusalem on July 24, 2020. (Photo by Ammar Awad/Reuters)

Palestinian Anoush Belian poses on stage during a bodybuilding competition in Tel Aviv, Israel August 22, 2015. Belian, from Jerusalem, on Saturday became the first Palestinian female bodybuilder to participate and win an Israeli competition. Belian won the Miss Fitness category in the competition organised by the National Amateur Bodybuilders Association (NABBA) in Israel. (Photo by Ammar Awad/Reuters)