Winner. “I took this vertical image in the Quarry Bay district of Hong Kong during the dusk ‘blue hour’, when there was a perfect balance between the ambient light in the sky and the artificial lights of the high-rise residential buildings”. (Photo by Jatinder Heer/The Guardian)

Behaviour winner; Courting Royals: two royal terns in courtship display by Kristian Bell. Another beautiful morning on a beautiful beach on the Gulf Coast of Florida seemed to prompt these two royal terns to commence an intricate courtship dance. The photograph was taken with a Canon 300mm lens and 2x extender. (Photo by Kristian Bell/Deakin University/Royal Society Publishing Photography Competition 2018)

Honourable mention, Behaviour category. Toss the scorpion – Indian roller playing with its kill by Susmita Datta. The image was taken during an early morning safari drive at Tadoba Andhari Tiger Reserve in India. (Photo by Susmita Datta/PA Wire/Royal Society Publishing Photography Competition 2017)

A woman wearing a scarf to cover her face looks on as she waits for a passenger bus on a smoggy morning in New Delhi, India, November 8, 2017. The Great Smog of Delhi is an ongoing severe air-pollution event in New Delhi and adjoining areas in the National Capital Territory of India, in November 2017. Air pollution at this time peaked on both PM 2.5 and PM 10 levels. It has been reported as one of the worst levels of air quality in Delhi since 1999. (Photo by Saumya Khandelwal/Reuters)

The 2019 Nissan Altima sits covered before being presented at the New York Auto Show in the Manhattan borough of New York City, New York, U.S., March 28, 2018. (Photo by Brendan McDermid/Reuters)

A participant covers himself with a blanket as it rains at the venue of the World Culture Festival on the banks of a river in New Delhi, India, March 11, 2016. (Photo by Adnan Abidi/Reuters)

Historical reenactor Troy Kotz poses with a mug of wassail in the parlor of the Guyon-Lake-Tysen House during the annual Candlelight Tour of Historic Richmond Town, Saturday, December 13, 2014, in the Staten Island borough of New York. The annual event lets visitors experience an authentic Victorian Christmas. This image was made using the TinType by Hipstamatic iPhone app. (Photo by Chad Rachman/New York Post)

Swimmers in fancy dress splash as they participate in the New Year's Day Loony Dook swim at South Queensferry, Scotland January 1, 2015. (Photo by Russell Cheyne/Reuters)