Winner – Animal Portraits: The pose by Mogens Trolle, Denmark. A young male proboscis monkey cocks his head slightly and closes his eyes. Unexpected pale blue eyelids now complement his immaculately groomed auburn hair. He poses for a few seconds as if in meditation. He is a wild visitor to the feeding station at Labuk Bay Proboscis Monkey Sanctuary in Sabah, Borneo – “the most laid-back character”, says Trolle, “quite unlike anything I’ve ever seen on another monkey” – connects us, he hopes, with a fellow primate. (Photo by Mogens Trolle/Wildlife Photographer of the Year 2020)

A tale of two foxes. Surprising behaviour, witnessed in Wapusk national park, on Hudson Bay, Canada, in early winter. Red foxes don’t actively hunt Arctic foxes, but where the ranges of two predators overlap, there can be conflict. Though the light was poor, the snow-covered tundra provided the backdrop for the moment that the red fox paused with the smaller fox in its mouth in a grim pose. (Photo by Don Gutoski/2015 Wildlife Photographer of the Year)

Arctic treasure by Sergey Gorshkov (Russia). An arctic fox carries its egg trophy from a raid on a snow goose nest and heads for a suitable burial spot. Finalist 2017, Animal Portraits. (Photo by Sergey Gorshkov/2017 Wildlife Photographer of the Year)

Komodo Judo by Andrey Gudkov. Finalist, Amphibians & Reptiles. Two large male Komodo dragons hissing angrily at each other in Indonesia’s Komodo national park. Komodo dragons can grow up to 8ft. (Photo by Andrey Gudkov/Wildlife Photographer of the Year 2015)

Andrew Parkinson, animal behaviour category winner: Crepuscular Contentment, Derbyshire. “In 15 years of working with badgers I’ve never seen a badger sit out in the open to have a scratch. I was sat concealed behind a tree and downwind so it was especially nice that the badger had his back to me, demonstrating just how inconspicuous and inconsequential my presence was”. (Photo by Andrew Parkinson/British Wildlife Photography Awards 2017)

Shooting star by Tony Wu, US/Japan. Winner, underwater. The electrifying reproductive dance of a giant sea star. As the surrounding water filled with sperm and eggs from spawning sea stars, Wu faced several challenges. Stuck in a small, enclosed bay with only a macro lens for photographing small subjects, he backed up to squeeze the undulating sea star into his field of view. The dancing posture of spawning sea stars rising and swaying may help release eggs and sperm, or may help sweep the eggs and sperm into the currents where they fertilise together in the water. Kinko Bay, Japan. (Photo by Tony Wu/Wildlife Photographer of the Year)

A golden huddle by Minqiang Lu, China. Two females and a male golden snub-nosed monkey huddle together to keep warm in the extreme cold. Threatened by forest loss and fragmentation, this endangered species is confined to central China. Restricted to living high up in the temperate forests, these monkeys – here in the Qinling mountains in Shaanxi province – feed mostly in the trees, on leaves, bark, buds and lichen. In heavy wind and snow, Minqiang walked up the mountain carrying his equipment. He stayed for half an hour in temperatures of –10C opposite the tree where the group was huddled before achieving this eye-level composition. (Photo by Minqiang Lu/Wildlife Photographer of the Year)

The British Wildlife Photography Awards winners have been revealed, with Lee Acaster from Suffolk taking home the top prize for his shot of a Graylag Goose in London. Acaster, who received £5,000, photographed the animal against an ominous London skyline, with The Shard clearly visible in the background. Here: “Urban Tourist (Graylag Goose)”. Urban category and overall winner. (Photo by Lee Acaster/British Wildlife Photography Awards 2014)