Guardian of the Mangroves – Overall Winner. Tanya Houppermans, Cuba. A curious American crocodile (Crocodylus acutus) swims right up to Tanya, at Gardens of the Queen (Jardines De La Reina), an archipelago off the coast of Cuba. It has been strictly protected since 1996, and is one of the most untouched marine ecosystems in the world. ‍ “The healthy population of American crocodiles is down to the pristine condition of the mangroves and I wanted to capture close ups of this gentle giant in its natural habitat. I hope this image can illustrate that protecting areas like this is so critical”. (Photo by Tanya Griffin Houppermans/Mangrove Photographer of the Year)

Youth category winner: Symphony No 5 by Giacomo Redaelli (Italy). (Photo by Giacomo Redaelli/2019 Nature Photographer of the Year)

“Eye of a toad”. Animal Portraits, Łukasz Bożycki, Poland. Early spring sees a pond near Łukasz’s home city of Warsaw, Poland, full of mating frogs and a few toads. On this March day, Łukasz shared the pond with them for an evening, sitting in the icy water in his chest-high waders, keeping as still as possible, despite the numbing cold, so that the amphibians could get used to him. “I wanted to find a fresh way of portraying the amphibians”, he says, “at water level”. Using a telephoto lens, he focused on one lone toad and waited for the sun to dip almost below the horizon before pressing the shutter, using flash to bring out the details in the shadow. His prize was “the glorious pool of sunset colour” and fiery glow of the toad’s eye. Nikon D80 + 70-300mm f4.5-5.6 lens + extension tube; 1/125 sec at f9 (-2.3 e/v); ISO 100; built-in flash. (Photo by Łukasz Bożycki)

Brad Snead walks past melting ice on the Cook Inlet beach, Saturday, March 26, 2011, in Kenai, Alaska, while looking for agates. (AP Photo/Peninsula Clarion, M. Scott Moon)

Actress Penelope Cruz attends a photocall promoting the film “Chromophobia” at the Palais during the 58th International Cannes Film Festival May 21, 2005 in Cannes, France. (Photo by M. J. Kim/Getty Images)

People board a train at the West Railway Station on January 7, 2012 in Beijing, China. China's railways are bracing themselves as millions of migrant labourers are set to leave cities for their native villages to celebrate the Chinese New Year. (Photo by Lintao Zhang/Getty Images)

Women take a selfie next to a police officer as they take part in New Year's Eve celebrations in the Bund, in Shanghai, China December 31, 2016. (Photo by Aly Song/Reuters)