Winner. “I took this vertical image in the Quarry Bay district of Hong Kong during the dusk ‘blue hour’, when there was a perfect balance between the ambient light in the sky and the artificial lights of the high-rise residential buildings”. (Photo by Jatinder Heer/The Guardian)

Behaviour winner; Courting Royals: two royal terns in courtship display by Kristian Bell. Another beautiful morning on a beautiful beach on the Gulf Coast of Florida seemed to prompt these two royal terns to commence an intricate courtship dance. The photograph was taken with a Canon 300mm lens and 2x extender. (Photo by Kristian Bell/Deakin University/Royal Society Publishing Photography Competition 2018)

Masked guests attend the “Grand Bal Christian Dior” during the Haute Couture Spring-Summer 2018 fashion collection presented in Paris, Monday, January 22, 2018. (Photo by Kamil Zihnioglu/AP Photo)

Filipino beauty candidates hide from the sun before a group photo during the media presentation of the Miss Philippines Earth 2017 held at a hotel in Manila, Philippines, 05 July 2017. A total of 40 environmentally aware and concerned Filipino young ladies will compete to represent the Philippines at the Miss Earth beauty pageant. Miss Earth is a pioneering beauty pageant that serves as a vehicle for environmental advocacy. (Photo by Mark R. Cristino/EPA)

A model walks the runway during the BLACK TAPE PROJECT show at New York Fashion Week Powered By Art Hearts Fashion at The Angel Orensanz Foundation on September 9, 2018 in New York City. (Photo by Arun Nevader/Getty Images for Art Hearts Fashion)

A model presents a creation during the 2018 Victoria's Secret Fashion Show in New York City, New York, U.S., November 8, 2018. (Photo by Mike Segar/Reuters)

A spectator waits for the start of the Jeremy Scott show during Fashion Week in New York, Thursday, February 8, 2018. (Photo by Andres Kudacki/AP Photo)

Runner-up. “The heavens opened and the streets around Shibuya, Tokyo, were suddenly drenched in even more colour and movement. I was taking photos (my friend was holding the umbrella) when I noticed this woman at the crossing. I’m pleased with the futuristic feel of the image”. (Photo by Katherine Bridgestock/The Guardian)