Behaviour winner; Courting Royals: two royal terns in courtship display by Kristian Bell. Another beautiful morning on a beautiful beach on the Gulf Coast of Florida seemed to prompt these two royal terns to commence an intricate courtship dance. The photograph was taken with a Canon 300mm lens and 2x extender. (Photo by Kristian Bell/Deakin University/Royal Society Publishing Photography Competition 2018)

Miss Colombia Ariadna Gutierrez, right, and Miss Philippines Pia Alonzo Wurtzbach wait for the announcer to name the new Miss Universe at the Miss Universe pageant on Sunday, December 20, 2015, in Las Vegas. Gutierrez was incorrectly crowned before Wurtzbach was named Miss Universe. (Photo by John Locher/AP Photo)

A contestant prepares at the backstage before the Miss Transqueen India 2018 transgender beauty pageant in Mumbai, India on October 7, 2018. (Photo by Francis Mascarenhas/Reuters)

Miss India Manushi Chhilar (C) reacts as she wins the 67 th Miss World contest final next to France Aurore Andrée Raphaëlle Kichenin (R) and Miss Kenya Magline Jeruto (L) in Sanya, on the tropical Chinese island of Hainan on November 18, 2017. (Photo by Nicolas Asfouri/AFP Photo)

Lebanese women smoke nargileh (water pipe) and watch a streaming broadcast of the FIFA World Cup 2022 group G soccer match between Brazil and Switzerland, at a café-restaurant in the area of Sabtiyeh, north of Beirut, on November 28, 2022. Lebanon being hit hard by the economic crisis, the country did not obtain the rights to broadcast the 2022 World Cup this year, depriving Lebanese fans of the World Cup. (Photo by Joseph Eid/AFP Photo)

Honourable mention, Behaviour category. Toss the scorpion – Indian roller playing with its kill by Susmita Datta. The image was taken during an early morning safari drive at Tadoba Andhari Tiger Reserve in India. (Photo by Susmita Datta/PA Wire/Royal Society Publishing Photography Competition 2017)

Youth category winner: Symphony No 5 by Giacomo Redaelli (Italy). (Photo by Giacomo Redaelli/2019 Nature Photographer of the Year)

Highly commended, mammals: Gelada after the storm – Marco Gaiotti (Italy). “Gelada baboons are the only monkey species in the world that feed on grasses. They are native to the tableland of Ethiopia. Every morning large family groups wander from their sleeping places in the steep rock face, up to 1,000 metres high, to the feeding grounds at the tablelands. This image clearly depicts their feeding strategy: they pull out bunches of grass, sort the stalks and then lift them to their mouth. This shot was taken towards the end of the rainy season after a heavy storm”. (Photo by Marco Gaiotti/2019 GDT European Wildlife Photographer of the Year)