A golden huddle by Minqiang Lu, China. Two females and a male golden snub-nosed monkey huddle together to keep warm in the extreme cold. Threatened by forest loss and fragmentation, this endangered species is confined to central China. Restricted to living high up in the temperate forests, these monkeys – here in the Qinling mountains in Shaanxi province – feed mostly in the trees, on leaves, bark, buds and lichen. In heavy wind and snow, Minqiang walked up the mountain carrying his equipment. He stayed for half an hour in temperatures of –10C opposite the tree where the group was huddled before achieving this eye-level composition. (Photo by Minqiang Lu/Wildlife Photographer of the Year)

Arachnids category 3rd place: dancing spider by Raed Ammari. Male jumping spiders (in this case a Phidippus insignarius) perform a courtship dance in which they almost form a heart shape with their legs. This one was shot in Colorado. (Photo by Raed Ammari/Luminar Bug Photographer of the Year 2020)

Tropical acrobatics by Adrià López Baucells in Manaus, Brazil. An unidentified South American marsupial, although the characteristic black markings on its face indicate it may be a mouse opossum. These small creatures are nocturnal and feed on bugs, fruit and bird eggs. (Photo by Adrià López Baucells/2019 Royal Society of Biology Photography Competition)

A) 1st place WINNER – Roy Rimmer. “This rat was in an outdoor set I made, the set up is two meters long and a meter wide made of Perspex,it has a plywood front with holes cut in for my camera and flash guns, I placed two rusty paint cans in the set up and the rat would leap from one can too the other, I had to use flash to freeze the action”.

Nosy neighbour by Sam Hobson, UK. Sam knew exactly who to expect when he set his camera on the wall one summer’s evening in a suburban street in Bristol, the UK’s famous fox city. He wanted to capture the inquisitive nature of the urban red fox in a way that would pique the curiosity of its human neighbours about the wildlife around them. (Photo by Sam Hobson/2016 Wildlife Photographer of the Year)

M42 Subtle V1 cropped. One of the most well-known astronomical objects in our universe is the Orion Nebula and this image depicts the wider region of the Orion Molecular Cloud Complex that is its home. This complex also includes another popular target for astrophotographers, the Horsehead Nebula, as well as Barnard’s Loop and the Running Man Nebula, which can be seen to the left of this photograph. (Photo by Patrick Gilliland)

Highly commended birds : Crested guan by Tim Hunt (UK). ‘This photo shows a crested guan in the cloud forests of Costa Rica as it pauses while foraging on a lone branch. Due to the clouds that are so typical for this habitat, I could often only photograph the bird’s silhouette against a grey sky. But then, for a short moment only, the sun broke through the clouds, and I overexposed the image by over two stops in order to blow out the background and allow this beautifully marked bird to stand out’. (Photo by Tim Hunt/GDT European Wildlife Photographer of the Year 2015)

Behaviour: Mammals category. Giant Gathering by Tony Wu, USA. Dozens of sperm whales mingled noisily off Sri Lanka’s northeast coast, stacked as far down as Tony could see. This was a congregation of dozens of social units, like a gathering of the clans. Aggregations like this could be a critical part of the whales’ rich social lives but are rarely reported. Some two thirds of the population was wiped out before commercial whaling was banned in 1986. This kind of major gathering could be “a sign that populations are recovering”, says Tony. (Photo by Tony Wu/Wildlife Photographer of the Year 2017)