Looking for love by Tony Wu, USA. Highly commended, Animal Portraits. “Accentuating his mature appearance with pastel colours, protruding lips and an outstanding pink forehead, this Asian sheepshead wrasse sets out to impress females and see off rivals, which he will head-butt and bite, near Japan’s remote Sado Island. Individuals start out as females, and when they reach a certain age and size – up to a metre (more than 3 feet) long – can transform into males. Long-lived and slow-growing, the species is intrinsically vulnerable to overfishing”. (Photo by Tony Wu/2018 Wildlife Photographer of the Year)

Shortlisted images in the running to be crowned Photobox Instagram photograph of the year range from furry friends to the Holi festival to the meaning of love. Judges, including the Guardian’s former picture editor Eamonn McCabe, have whittled down 180,000 submissions to unearth a shortlist that celebrates the best of social media. Winners will be announced October 3, 2018. Here: Najac, France. Travel shortlist. (Photo by Aaron Jenkin/@aaronjenkin)

“Tasty Nature” by Dipesh Bhatt. “The blue banded bee (Amegilla cingulata) is pointing this sheath towards the flowers in preparation for sipping nectar”. Taken in Gujarat, India. (Photo by Dipesh Bhatt/2016 Royal Society of Biology Photographer of the Year competition)

Wide-angle category winner. Part of the Illusion by Marcus Blatchford (UK). Location: National Dive & Activity Centre, Chepstow, Wales. “This is the deepest inland dive centre in the UK. The dive plan was to explore the deep end, but this time I dived ‘unplugged’ (without my strobes). With the exception of this change to my camera technique, there were no planned shots I wanted to achieve – just a fun dive with ad-hoc photos along the way. Shortly after this photo was captured, in 6C water and two hours of decompression ahead of us, we turned and started the long ascent back to the surface”. (Photo by Marcus Blatchford/Underwater Photographer of the Year 2016)

Aurorae category runner-up: Lone Tree under a Scandinavian Aurora by Tom Archer (UK). The photographer decided to explore the area around the hotel on a very crisp -35C evening in Finnish Lapland. When he found this tree, he decided to wait for the misty conditions to change and could not believe his luck when the sky cleared and the aurora came out in the perfect spot. Archer spent about an hour photographing it before his camera started to lock up because of the harsh conditions, but by then he was happy to call it a night. (Photo by Tom Archer/2020 Astronomy Photographer of the Year)

Shortlisted: “Two big eyes” by Miao Yong (Zejiang province, China). Damselflies look over the leaves. “I was photographing insects in a park near my home when suddenly I found two damselflies in the grass. They kept flying and it was very difficult to focus until suddenly they parked behind a leaf”. (Photo by Miao Yong/2017 Royal Society of Biology Photographer of the Year)

British underwater photographer of the year – winner. “Love Birds” by Grant Thomas (UK). Location: Luss Pier, Loch Lomond, Scotland. Thomas’s initial idea was to frame a split shot of one swan feeding below the surface of the water but when he noticed how comfortable they were around him he was confident, with some patience, he could get that magical shot of the two. (Photo by Grant Thomas/UPY 2018)

“Red Arrows” at the 2016 royal air tattoo, RAF Fairford, UK. By Jonny Williams, finalist: action. (Photo by Jonny Williams/UK National Geographic Traveller Photography Competition 2018)