With its huge eyes, comical name and diminutive size, Mark R. Smith’s image of a baby Hawaiian bobtail squid can’t help but raise a smile. A curiously endearing creature, the cephalopod is just 1.5cm across, its mantle cavity bearing more than a passing resemblance to a rather natty shower cap. But it is also a beautiful example of symbiosis – nature’s version of “I’ll scratch your back if you scratch mine” – for on the underside of the squid is a light organ which houses bioluminescent bacteria. The squid offers the bacteria protection and food, while the bacteria emit a glow – a handy trait that the squid uses to offset its silhouette, helping it to evade predators in the depths below. Mark R. Smith’s entry combines several images of a Hawaiian bobtail squid with different focus lengths to create a final picture with greater depth of field than normal. (Photo by Mark R. Smith/Wellcome Images/Macroscopic Solutions)

Andrew Parkinson, animal behaviour category winner: Crepuscular Contentment, Derbyshire. “In 15 years of working with badgers I’ve never seen a badger sit out in the open to have a scratch. I was sat concealed behind a tree and downwind so it was especially nice that the badger had his back to me, demonstrating just how inconspicuous and inconsequential my presence was”. (Photo by Andrew Parkinson/British Wildlife Photography Awards 2017)