A frog appears to have a big smile for the camera, Russia. Things are heating up at the Comedy Wildlife Photography Awards as the shortlisted final 40 entries are revealed. This year's competition has featured over 2200 hilarious entries from around the world with photos including a fox face planting in the snow, a dancing brown bear and a pair of seemingly headless penguins – all beautifully photographed with perfect comedy timing and a strong conservation message. (Photo by Artyom Krivosheev/Barcroft Images/Comedy Wildlife Photography Awards 2016)

“Family disagreement”. (Photo by Vlado Pirsa/Comedy Wildlife Photo Awards 2019)

Hidden Britain category winner. Garden Spider by Alan Smith from Reading, Berkshire. (Photo by Alan Smith/British Wildlife Photography Awards/PA Wire Press Association)

The British Wildlife Photography Awards winners have been revealed, with Lee Acaster from Suffolk taking home the top prize for his shot of a Graylag Goose in London. Acaster, who received £5,000, photographed the animal against an ominous London skyline, with The Shard clearly visible in the background. Here: “Urban Tourist (Graylag Goose)”. Urban category and overall winner. (Photo by Lee Acaster/British Wildlife Photography Awards 2014)

Breath of an Arctic fox by Marco Gaiotti, Italy. Marco was watching this little Arctic fox as it incessantly called another nearby. Gradually he noticed the fox’s wet breath was quickly freezing in the air after each call. It was late winter in Spitsbergen, Svalbard, and the air was -35C (-31F). Photographing Arctic foxes is often frustrating, as they are normally running around fast in search of food, but this one was very relaxed and let Marco get close enough to focus on it, with the light glowing perfectly in the background. (Photo by Marco Gaiotti/Wildlife Photographer of the Year 2021)

“Romantic”. A brown bear in Martinselkonen, Finland. (Photo by Valtteri Mulkahainen/Comedy Wildlife Photography Awards)

A golden huddle by Minqiang Lu, China. Two females and a male golden snub-nosed monkey huddle together to keep warm in the extreme cold. Threatened by forest loss and fragmentation, this endangered species is confined to central China. Restricted to living high up in the temperate forests, these monkeys – here in the Qinling mountains in Shaanxi province – feed mostly in the trees, on leaves, bark, buds and lichen. In heavy wind and snow, Minqiang walked up the mountain carrying his equipment. He stayed for half an hour in temperatures of –10C opposite the tree where the group was huddled before achieving this eye-level composition. (Photo by Minqiang Lu/Wildlife Photographer of the Year)

This year’s overall winner and winner of the coast and marine category is George Stoyle with his image “Hitchhikers” of a Lion’s mane jellyfish, photographed at St Kilda, off the Island of Hirta, Scotland. (Photo by George Stoyle/British Wildlife Photography Awards 2016)