Breath of an Arctic fox by Marco Gaiotti, Italy. Marco was watching this little Arctic fox as it incessantly called another nearby. Gradually he noticed the fox’s wet breath was quickly freezing in the air after each call. It was late winter in Spitsbergen, Svalbard, and the air was -35C (-31F). Photographing Arctic foxes is often frustrating, as they are normally running around fast in search of food, but this one was very relaxed and let Marco get close enough to focus on it, with the light glowing perfectly in the background. (Photo by Marco Gaiotti/Wildlife Photographer of the Year 2021)

Looking for love by Tony Wu, USA. Highly commended, Animal Portraits. “Accentuating his mature appearance with pastel colours, protruding lips and an outstanding pink forehead, this Asian sheepshead wrasse sets out to impress females and see off rivals, which he will head-butt and bite, near Japan’s remote Sado Island. Individuals start out as females, and when they reach a certain age and size – up to a metre (more than 3 feet) long – can transform into males. Long-lived and slow-growing, the species is intrinsically vulnerable to overfishing”. (Photo by Tony Wu/2018 Wildlife Photographer of the Year)

A zebra walks through the recently flooded Safari Park located at Safari World November 2, 2011 in Bangkok, Thailand. (Photo by Paula Bronstein /Getty Images)

Behaviour: Mammals category. Giant Gathering by Tony Wu, USA. Dozens of sperm whales mingled noisily off Sri Lanka’s northeast coast, stacked as far down as Tony could see. This was a congregation of dozens of social units, like a gathering of the clans. Aggregations like this could be a critical part of the whales’ rich social lives but are rarely reported. Some two thirds of the population was wiped out before commercial whaling was banned in 1986. This kind of major gathering could be “a sign that populations are recovering”, says Tony. (Photo by Tony Wu/Wildlife Photographer of the Year 2017)

Nosy neighbour by Sam Hobson, UK. Sam knew exactly who to expect when he set his camera on the wall one summer’s evening in a suburban street in Bristol, the UK’s famous fox city. He wanted to capture the inquisitive nature of the urban red fox in a way that would pique the curiosity of its human neighbours about the wildlife around them. (Photo by Sam Hobson/2016 Wildlife Photographer of the Year)

This year’s overall winner and winner of the coast and marine category is George Stoyle with his image “Hitchhikers” of a Lion’s mane jellyfish, photographed at St Kilda, off the Island of Hirta, Scotland. (Photo by George Stoyle/British Wildlife Photography Awards 2016)

Highly commended birds : Crested guan by Tim Hunt (UK). ‘This photo shows a crested guan in the cloud forests of Costa Rica as it pauses while foraging on a lone branch. Due to the clouds that are so typical for this habitat, I could often only photograph the bird’s silhouette against a grey sky. But then, for a short moment only, the sun broke through the clouds, and I overexposed the image by over two stops in order to blow out the background and allow this beautifully marked bird to stand out’. (Photo by Tim Hunt/GDT European Wildlife Photographer of the Year 2015)