Respect, Kronotsky nature reserve, Russia. The photographer’s cat, Ryska – her name means little lynx in Russian – stands outside their cabin and with aggressive posturing warns off a fox. In winter, foxes would regularly visit the cabin searching for food. If one peered in at the window, possible when the snow was deep, Ryska would sit on the other side, fur raised, and growl. When outside, she would hold her ground. The foxes were not always frightened and so encounters could be a sort of dance. (Photo by Igor Shpilenok/Unforgettable Behaviour/NHM)

Brush fires in Everglades National Park, Florida make for a dramatic Milky Way in this panorama. A photographer set out to inspire awe in nature's wonders through the tropical wetlands of Florida. (Photo by Mark Andrew Thomas/Rex Feature/Shutterstock)

Birds behaviour winner: Land of the Eagle by Audun Rikardsen, Norway. High on a ledge, on the coast near his home in northern Norway, Rikardsen carefully positioned an old tree branch that he hoped would make a perfect golden eagle lookout. To this, he bolted a tripod head with a camera, flashes and motion sensor attached, and built himself a hide a short distance away. From time to time, he left road‑kill carrion nearby. Very gradually – over the next three years – a golden eagle got used to the camera and started to use the branch regularly to survey the coast below. (Photo by Audun Rikardsen/2019 Wildlife Photographer of the Year)

Sompob Sasi-Smit is a professional nature and wildlife photographer based in Thailand. The artist has created a fantastic portfolio capturing the natural beauty and the charm of wild animals mostly among national parks.

The National Geographic Photo Ark is a travelling exhibition of photographer Joel Sartore’s quest to create a photo archive of biodiversity around the world. So far, Sartore has captured studio portraits of more than 6,000 species – a number that he hopes to double. On 1 July, the ark will open at Melbourne zoo – the first time it has been exhibited in the southern hemisphere. More than 50 portraits will be on display, including many of Australian endangered animals being protected by programs at the zoo itself. These captions have been edited from text supplied by Melbourne zoo. Here: Barking owl. (Photo by Joel Sartore/National Geographic Photo Ark/The Guardian)

A lone male cheetah is set upon by a pack of African wild dogs. Peter Haygarth had been following the dogs as they hunted in Zimanga Private Game Reserve in KwaZulu-Natal, South Africa. On first encountering the cheetah, the dogs were wary, but as the rest of the pack arrived, their confidence grew and they began to encircle the cat. Peter kept his focus on the cat’s face. In a few minutes the spat was over as the cheetah fled. (Behaviour: mammals category). (Photo by Peter Haygarth)

More than 1,500 snappers submitted their most hilarious pictures of all creatures great and small, and now 45 have made the cut. From drunken-eyed owls to embarrassed chipmunks and laughing goats – the finalists in the Comedy Wildlife Photography Awards are guaranteed to raise a smile. Here: 'Dancing sifaka'. (Photo by Alison Buttigieg/Comedy Wildlife Photography Awards/Mercury Press)

“Eye of a toad”. Animal Portraits, Łukasz Bożycki, Poland. Early spring sees a pond near Łukasz’s home city of Warsaw, Poland, full of mating frogs and a few toads. On this March day, Łukasz shared the pond with them for an evening, sitting in the icy water in his chest-high waders, keeping as still as possible, despite the numbing cold, so that the amphibians could get used to him. “I wanted to find a fresh way of portraying the amphibians”, he says, “at water level”. Using a telephoto lens, he focused on one lone toad and waited for the sun to dip almost below the horizon before pressing the shutter, using flash to bring out the details in the shadow. His prize was “the glorious pool of sunset colour” and fiery glow of the toad’s eye. Nikon D80 + 70-300mm f4.5-5.6 lens + extension tube; 1/125 sec at f9 (-2.3 e/v); ISO 100; built-in flash. (Photo by Łukasz Bożycki)