A golden huddle by Minqiang Lu, China. Two females and a male golden snub-nosed monkey huddle together to keep warm in the extreme cold. Threatened by forest loss and fragmentation, this endangered species is confined to central China. Restricted to living high up in the temperate forests, these monkeys – here in the Qinling mountains in Shaanxi province – feed mostly in the trees, on leaves, bark, buds and lichen. In heavy wind and snow, Minqiang walked up the mountain carrying his equipment. He stayed for half an hour in temperatures of –10C opposite the tree where the group was huddled before achieving this eye-level composition. (Photo by Minqiang Lu/Wildlife Photographer of the Year)

Shooting star by Tony Wu, US/Japan. Winner, underwater. The electrifying reproductive dance of a giant sea star. As the surrounding water filled with sperm and eggs from spawning sea stars, Wu faced several challenges. Stuck in a small, enclosed bay with only a macro lens for photographing small subjects, he backed up to squeeze the undulating sea star into his field of view. The dancing posture of spawning sea stars rising and swaying may help release eggs and sperm, or may help sweep the eggs and sperm into the currents where they fertilise together in the water. Kinko Bay, Japan. (Photo by Tony Wu/Wildlife Photographer of the Year)

Polar frame, by Dmitry Kokh, Russia. When Kokh’s boat approached the small island of Kolyuchin in the Russian High Arctic, which had been abandoned by humans since 1992, he was surprised to spot movement in one of the houses. Binoculars revealed polar bears – more than 20 in total – exploring the ghost town. Dmitry used a low-noise drone to document them. (Photo by Dmitry Kokh/Wildlife Photographer of the Year 2022)

Breath of an Arctic fox by Marco Gaiotti, Italy. Marco was watching this little Arctic fox as it incessantly called another nearby. Gradually he noticed the fox’s wet breath was quickly freezing in the air after each call. It was late winter in Spitsbergen, Svalbard, and the air was -35C (-31F). Photographing Arctic foxes is often frustrating, as they are normally running around fast in search of food, but this one was very relaxed and let Marco get close enough to focus on it, with the light glowing perfectly in the background. (Photo by Marco Gaiotti/Wildlife Photographer of the Year 2021)

Winner, photojournalism. Elephant in the room, by Adam Oswell, Australia Zoo. Visitors watch a young elephant performing underwater. Oswell was disturbed by this scene, and organisations concerned with the welfare of captive elephants say performances like this encourage unnatural behaviour. In Thailand, there are now more elephants in captivity than in the wild. With the Covid pandemic causing tourism to collapse, elephant sanctuaries are becoming overwhelmed with animals that can no longer be looked after by their owners. (Photo by Adam Oswell/Wildlife Photographer of the Year 2021)

Winner – Animal Portraits: The pose by Mogens Trolle, Denmark. A young male proboscis monkey cocks his head slightly and closes his eyes. Unexpected pale blue eyelids now complement his immaculately groomed auburn hair. He poses for a few seconds as if in meditation. He is a wild visitor to the feeding station at Labuk Bay Proboscis Monkey Sanctuary in Sabah, Borneo – “the most laid-back character”, says Trolle, “quite unlike anything I’ve ever seen on another monkey” – connects us, he hopes, with a fellow primate. (Photo by Mogens Trolle/Wildlife Photographer of the Year 2020)

Highly commended, mammals: Gelada after the storm – Marco Gaiotti (Italy). “Gelada baboons are the only monkey species in the world that feed on grasses. They are native to the tableland of Ethiopia. Every morning large family groups wander from their sleeping places in the steep rock face, up to 1,000 metres high, to the feeding grounds at the tablelands. This image clearly depicts their feeding strategy: they pull out bunches of grass, sort the stalks and then lift them to their mouth. This shot was taken towards the end of the rainy season after a heavy storm”. (Photo by Marco Gaiotti/2019 GDT European Wildlife Photographer of the Year)

Birds behaviour winner: Land of the Eagle by Audun Rikardsen, Norway. High on a ledge, on the coast near his home in northern Norway, Rikardsen carefully positioned an old tree branch that he hoped would make a perfect golden eagle lookout. To this, he bolted a tripod head with a camera, flashes and motion sensor attached, and built himself a hide a short distance away. From time to time, he left road‑kill carrion nearby. Very gradually – over the next three years – a golden eagle got used to the camera and started to use the branch regularly to survey the coast below. (Photo by Audun Rikardsen/2019 Wildlife Photographer of the Year)