Wakodahatchee wetlands, Delray Beach, Florida, US. Equipped with sinewy necks and spear-like bills, great blue herons can lunge with fearsome speed to strike their aquatic prey. Adults will also employ rapid stabbing motions as one aspect of their complex courtship displays; they’re seemingly dangerous moves, but fitting to the intensity of mating season. (Photo by Melissa Rowell/Audubon Photography Awards)

Highly commended, mammals: Gelada after the storm – Marco Gaiotti (Italy). “Gelada baboons are the only monkey species in the world that feed on grasses. They are native to the tableland of Ethiopia. Every morning large family groups wander from their sleeping places in the steep rock face, up to 1,000 metres high, to the feeding grounds at the tablelands. This image clearly depicts their feeding strategy: they pull out bunches of grass, sort the stalks and then lift them to their mouth. This shot was taken towards the end of the rainy season after a heavy storm”. (Photo by Marco Gaiotti/2019 GDT European Wildlife Photographer of the Year)

Shooting star by Tony Wu, US/Japan. Winner, underwater. The electrifying reproductive dance of a giant sea star. As the surrounding water filled with sperm and eggs from spawning sea stars, Wu faced several challenges. Stuck in a small, enclosed bay with only a macro lens for photographing small subjects, he backed up to squeeze the undulating sea star into his field of view. The dancing posture of spawning sea stars rising and swaying may help release eggs and sperm, or may help sweep the eggs and sperm into the currents where they fertilise together in the water. Kinko Bay, Japan. (Photo by Tony Wu/Wildlife Photographer of the Year)

Yala national park, Sri Lanka. Category: Action. “The daylight was fading and I was scanning the bush for a leopard. Then, I saw this garden lizard and a common bronzeback snake in the road. When the snake struck, the lizard performed amazing acrobatics to avoid it. The snake didn’t enjoy the extra attention from the crowd of jeeps. The distraction gave the lizard the chance to live another day”. (Photo by Sajith Buddikha Withanage/National Geographic Traveller UK)

Looking for love by Tony Wu, USA. Highly commended, Animal Portraits. “Accentuating his mature appearance with pastel colours, protruding lips and an outstanding pink forehead, this Asian sheepshead wrasse sets out to impress females and see off rivals, which he will head-butt and bite, near Japan’s remote Sado Island. Individuals start out as females, and when they reach a certain age and size – up to a metre (more than 3 feet) long – can transform into males. Long-lived and slow-growing, the species is intrinsically vulnerable to overfishing”. (Photo by Tony Wu/2018 Wildlife Photographer of the Year)

Actor Bella Thorne (L) and model Amber Rose at the NYLON Young Hollywood Party at AVENUE Los Angeles on May 2, 2017 in Los Angeles, California. (Photo by Joe Scarnici/Getty Images for NYLON)

The competition, which is run by the Royal Observatory Greenwich in association with BBC Sky at Night Magazine, is now in its sixth year and has received over 2,500 entries. Photo: Centre of the Heart Nebula by Ivan Eder (Hungary). Situated 7500 light years away in the “W”-shaped constellation of Cassiopeia, the Heart Nebula is a vast region of glowing gas, energized by a cluster of young stars at its centre. The image depicts the central region, where dust clouds are being eroded and moulded into rugged shapes by the searing cosmic radiation. (Photo by Ivan Eder)

British Waters Wide Angle category runner-up. Grass snake swimming along a garden pond by Jack Perks (UK)in Nottinghamshire, UK. “I’m always on the lookout for unusual freshwater subjects and grass snakes are a species I’ve been after for years. I was told about a pond where the odd grass snake hangs around the lilly pads for frogs. I put my drysuit on and got into the water and could see one slithering along the surface. Slowly making my way towards it with my head only just poking above I got the spilt shot”. (Photo by Jack Perks/Underwater Photographer of the Year 2019)