Winner. “I took this vertical image in the Quarry Bay district of Hong Kong during the dusk ‘blue hour’, when there was a perfect balance between the ambient light in the sky and the artificial lights of the high-rise residential buildings”. (Photo by Jatinder Heer/The Guardian)

Behaviour winner; Courting Royals: two royal terns in courtship display by Kristian Bell. Another beautiful morning on a beautiful beach on the Gulf Coast of Florida seemed to prompt these two royal terns to commence an intricate courtship dance. The photograph was taken with a Canon 300mm lens and 2x extender. (Photo by Kristian Bell/Deakin University/Royal Society Publishing Photography Competition 2018)

Runner-up. “The heavens opened and the streets around Shibuya, Tokyo, were suddenly drenched in even more colour and movement. I was taking photos (my friend was holding the umbrella) when I noticed this woman at the crossing. I’m pleased with the futuristic feel of the image”. (Photo by Katherine Bridgestock/The Guardian)

Elena Chernyshova's vision of Norilsk, Russia, the northernmost city in the world, is a series of surprises by which she extracts otherworldly beauty from ugly realities. Here: 2013. A woman is visible through a narrow passageway between two buildings. Norilsk's urban spaces were designed to shorten distances around large developments and give residents maximum protection from arctic winds. (Photo by Elena Chernyshova)

Doris Pennman adjusts her umbrella as she walks along Mission Beach on January 8, 2005 in San Diego, Califonia. (Photo by Sandy Huffaker/Getty Images)

Members of an American landing party assist troops whose landing craft was sunk by enemy fire off Omaha beach, near Colleville sur Mer, France, June 6, 1944. REUTERS/Weintraub/US National Archives

People immerse themselves in an ice hole in the Chulym river, with the air temperature at about minus 12 degrees Celsius (10.4 degrees Fahrenheit), during celebrations for the Russian Orthodox Epiphany outside of the town of Nazarovo, Krasnoyarsk region January 19, 2015. (Photo by Ilya Naymushin/Reuters)

Members of a historical re-enactment group dressed as U.S. and Soviet Army soldiers take part in Elbe Day celebrations, in eastern German city of Torgau at the river Elbe, April 25, 2015. Elbe Day commemorates the encounter of the Allies 70 years ago, on the 25th of April 1945 when American and Soviet army units joined together on the destroyed bridge over the river Elbe. The photograph of the meaningful handshake made its way around the world and became a symbol of the near end of World War II in Europe. (Photo by Stefanie Loos/Reuters)