Shooting star by Tony Wu, US/Japan. Winner, underwater. The electrifying reproductive dance of a giant sea star. As the surrounding water filled with sperm and eggs from spawning sea stars, Wu faced several challenges. Stuck in a small, enclosed bay with only a macro lens for photographing small subjects, he backed up to squeeze the undulating sea star into his field of view. The dancing posture of spawning sea stars rising and swaying may help release eggs and sperm, or may help sweep the eggs and sperm into the currents where they fertilise together in the water. Kinko Bay, Japan. (Photo by Tony Wu/Wildlife Photographer of the Year)

A huge atlas moth, with a wingspan of more than 9in, photographed on an areca nut plantation in Sirsi, India. (Photo by Uday Hegde/Close Up Photographer of the Year)

A pool ingeniously filled with clear water in the middle of Lake Kerniki in Greece enabled this shot of feeding pelicans. (Photo by Bence Mate/Close Up Photographer of the Year 2020)

First place, The Beauty of Plants. “There are many stages of lotus growth on display at the Aquatic Gardens, but to come across twotwisted dancing stems of nelumbo nucifera was unexpected and quite magical”. (Photo by Kathleen Furey/The Guardian)

“Horst Faas (28 April 1933 – 10 May 2012) was a German photo-journalist and two-time Pulitzer Prize winner. He is best-known for his images of the Vietnam War”. – Wikipedia

Photo: A U.S. crewman runs from a crashed CH-21 Shawnee troop helicopter near the village of Ca Mau in the southern tip of South Vietnam, December 11, 1962. Two helicopters crashed without serious injuries during a government raid on the Viet Cong-infiltrated area. Both helicopters were destroyed to keep them out of enemy hands. (Photo by AP Photo/Horst Faas)

Commended. In late May, about a quarter of a million snow geese arrive from North America to nest on Wrangel Island, in northeastern Russia. They form the world's largest breeding colony of snow geese. Photographer Sergey Gorshkov spent two months on the remote island photographing the unfolding dramas. Arctic foxes take advantage of the abundance of eggs, caching surplus eggs for leaner times. But a goose (here the gander) is easily a match for a fox, which must rely on speed and guile to steal eggs. “The battles were fairly equal”, notes Sergey, “and I only saw a fox succeed in grabbing an egg on a couple of occasions, despite many attempts”. Surprisingly, “the geese lacked any sense of community spirit”, he adds, “and never reacted when a fox harassed a neighboring pair nesting close by”. (Photo by Sergey Gorshkov/Veolia Environnement Wildlife Photographer)

The Invasion. A quiet street in Macau. Modernization around is quickly changing the city, as documented by Paul Tsui. (Photo by Paul Tsui/National Geographic Travel Photographer of the Year Contest)

Best portfolio winner. Roseate Spoonbill, Platalea ajaja, by Petr Bambousek, Czech Republic. (Photo by Petr Bambousek/2018 Bird Photographer of the Year)