Birds behaviour winner: Land of the Eagle by Audun Rikardsen, Norway. High on a ledge, on the coast near his home in northern Norway, Rikardsen carefully positioned an old tree branch that he hoped would make a perfect golden eagle lookout. To this, he bolted a tripod head with a camera, flashes and motion sensor attached, and built himself a hide a short distance away. From time to time, he left road‑kill carrion nearby. Very gradually – over the next three years – a golden eagle got used to the camera and started to use the branch regularly to survey the coast below. (Photo by Audun Rikardsen/2019 Wildlife Photographer of the Year)

The National Geographic Photo Ark is a travelling exhibition of photographer Joel Sartore’s quest to create a photo archive of biodiversity around the world. So far, Sartore has captured studio portraits of more than 6,000 species – a number that he hopes to double. On 1 July, the ark will open at Melbourne zoo – the first time it has been exhibited in the southern hemisphere. More than 50 portraits will be on display, including many of Australian endangered animals being protected by programs at the zoo itself. These captions have been edited from text supplied by Melbourne zoo. Here: Barking owl. (Photo by Joel Sartore/National Geographic Photo Ark/The Guardian)

A lone male cheetah is set upon by a pack of African wild dogs. Peter Haygarth had been following the dogs as they hunted in Zimanga Private Game Reserve in KwaZulu-Natal, South Africa. On first encountering the cheetah, the dogs were wary, but as the rest of the pack arrived, their confidence grew and they began to encircle the cat. Peter kept his focus on the cat’s face. In a few minutes the spat was over as the cheetah fled. (Behaviour: mammals category). (Photo by Peter Haygarth)

A tale of two foxes. Surprising behaviour, witnessed in Wapusk national park, on Hudson Bay, Canada, in early winter. Red foxes don’t actively hunt Arctic foxes, but where the ranges of two predators overlap, there can be conflict. Though the light was poor, the snow-covered tundra provided the backdrop for the moment that the red fox paused with the smaller fox in its mouth in a grim pose. (Photo by Don Gutoski/2015 Wildlife Photographer of the Year)

Arctic treasure by Sergey Gorshkov (Russia). An arctic fox carries its egg trophy from a raid on a snow goose nest and heads for a suitable burial spot. Finalist 2017, Animal Portraits. (Photo by Sergey Gorshkov/2017 Wildlife Photographer of the Year)

The winners of The London’s Natural History Museum's prestigious Wildlife Photographer of the Year for 2013 have finally been unveiled. Selected from almost 43,000 entries from 96 countries, the winners offer a glimpse of the stunning array of natural beauty on our planet. Photo: Canada: “Lucky pounce”. “Anticipating the pounce – that was the hardest part”, says Connor, who had come to Yellowstone National Park, Wyoming, USA, in search of wildlife as much as the spectacular landscape. He had found this fox, his first ever, on his last day in the park. It was so absorbed in hunting that Connor had plenty of time to get out of the car and settle behind a rock. It quartered the grassland, back and forth, and then started staring intently at a patch of ground, giving Connor just enough warning of the action to come. When it sprung up, Connor got his shot. And when it landed, the fox got his mouse. (Photo by Connor Stefanison/Wildlife Photographer of the Year 2013)

An alarmed squirrel. (Photo by Mary McGowan/Barcroft Images/Comedy Wildlife Photography Awards)

Bloodthirsty by Thomas P. Peschak, Germany/South Africa — winner, Behaviour: birds. When rations run short on Wolf Island, in the remote northern Galápagos, the sharp-beaked ground finches become vampires. Their sitting targets are Nazca boobies and other large birds. The finches rely on a scant diet of seeds and insects, which regularly dries up, so they drink blood to survive. ‘I’ve seen more than half a dozen finches drinking from a single Nazca booby,’ says Tom. Rather than leave their nests the boobies tolerate the vampires, and the blood loss doesn’t seem to cause permanent harm. (Photo by Thomas P. Peschak/2018 Wildlife Photographer of the Year)