First place, The Beauty of Plants. “There are many stages of lotus growth on display at the Aquatic Gardens, but to come across twotwisted dancing stems of nelumbo nucifera was unexpected and quite magical”. (Photo by Kathleen Furey/The Guardian)

Shooting star by Tony Wu, US/Japan. Winner, underwater. The electrifying reproductive dance of a giant sea star. As the surrounding water filled with sperm and eggs from spawning sea stars, Wu faced several challenges. Stuck in a small, enclosed bay with only a macro lens for photographing small subjects, he backed up to squeeze the undulating sea star into his field of view. The dancing posture of spawning sea stars rising and swaying may help release eggs and sperm, or may help sweep the eggs and sperm into the currents where they fertilise together in the water. Kinko Bay, Japan. (Photo by Tony Wu/Wildlife Photographer of the Year)

A huge atlas moth, with a wingspan of more than 9in, photographed on an areca nut plantation in Sirsi, India. (Photo by Uday Hegde/Close Up Photographer of the Year)

Wide-angle category winner. Part of the Illusion by Marcus Blatchford (UK). Location: National Dive & Activity Centre, Chepstow, Wales. “This is the deepest inland dive centre in the UK. The dive plan was to explore the deep end, but this time I dived ‘unplugged’ (without my strobes). With the exception of this change to my camera technique, there were no planned shots I wanted to achieve – just a fun dive with ad-hoc photos along the way. Shortly after this photo was captured, in 6C water and two hours of decompression ahead of us, we turned and started the long ascent back to the surface”. (Photo by Marcus Blatchford/Underwater Photographer of the Year 2016)

“Tasty Nature” by Dipesh Bhatt. “The blue banded bee (Amegilla cingulata) is pointing this sheath towards the flowers in preparation for sipping nectar”. Taken in Gujarat, India. (Photo by Dipesh Bhatt/2016 Royal Society of Biology Photographer of the Year competition)

Shortlisted: Pooyan Shadpoor, Houcheraghi. While walking along the shore of Larak, Iran – an island in the Persian Gulf – Shadpoor came across this luminous scene. The “magical lights of (the) plankton ... enchanted me so that I snapped the shot”, he writes. (Photo by Pooyan Shadpoor/2016 EPOTY)

Second place, Nature: double trapping. Picture taken in the Brazilian Pantanal ... when I downloaded the CF did not want to believe it ... The nature knows we always give magnificent events but sometimes extraordinary. (Photo by Massimiliano Bencivenni/National Geographic Travel Photographer of the Year Contest)

Australian photographer Brendan Fitzpatrick’s X-ray photographs expose the inner workings of toys. Fitzpatrick’s photographs are both whimsical and mechanical, evoking the curiosity of childhood and the desire to discover how things look and work from other perspectives. The strategic placement of wires, batteries, and screws are revealed, the complexity of the inside contrasting with the seemingly simplistic design of the outside. Fitzpatrick uses chest X-ray and mammogram machines to photograph flowers, toys, and creatures, then enhances the color in the images in order to more effectively distinguish the various parts that have been exposed. This photographs are part of series he calls “Invisible Light”. (Photo by Brendan Fitzpatrick)