Runner-up; Hon Yen marine ecosystem. Phu Yen, Vietnam. Every year between May and August, the coral of this rich and diverse ecosystem becomes exposed at low tide. (Photo by Truong Hoai Vu/Royal Society of Biology Photography Competition)

Winner – Animal Portraits: The pose by Mogens Trolle, Denmark. A young male proboscis monkey cocks his head slightly and closes his eyes. Unexpected pale blue eyelids now complement his immaculately groomed auburn hair. He poses for a few seconds as if in meditation. He is a wild visitor to the feeding station at Labuk Bay Proboscis Monkey Sanctuary in Sabah, Borneo – “the most laid-back character”, says Trolle, “quite unlike anything I’ve ever seen on another monkey” – connects us, he hopes, with a fellow primate. (Photo by Mogens Trolle/Wildlife Photographer of the Year 2020)

A sewer cleaner of Dhaka City Corporation cleaning out the city's sewers on May 03, 2017 in Dhaka, Bangladesh. Despite a rise in the number of deaths of manhole workers every year, workers regularly go into the manholes without any protective gear. (Photo by Zakir Chowdhury/Barcroft Images)

African migrants stranded in Costa Rica walk near garbage on the Inter-American highway at the border between Costa Rica and Nicaragua, in Penas Blancas, Costa Rica, September 8, 2016. (Photo by Juan Carlos Ulate/Reuters)

Male orangutan Percy is reflected in the Sekonyer River as he reaches over to try and touch a wooden klotok boat carrying crew and tourists, in Tanjung Puting National Park, in Kalimantan (Indonesian Borneo), Indonesia, September 4, 2013. Percy is a son, born and living in the wild, of a female orangutan named Princess, a freed former captive orangutan that anthropologist Dr Birute Galdikas rescued, rehabilitated and returned to the wild. (Photo by Barbara Walton/EPA)

Rebels of the Revolutionary Armed Forces of Colombia (FARC) visit as they wait to have lunch in El Diamante, Colombia, Friday, September 16, 2016. FARC rebels are gathering for a congress were delegates will debate and vote on the accord reached last month with the Colombian government to end five decades of war. (Photo by Ricardo Mazalan/AP Photo)

A child uses an umbrella to avoid rain falling on a fire at the northern Greek border point of Idomeni, Greece, Monday, April 25, 2016. Many thousands of migrants remain at the Greek border with Macedonia, hoping that the border crossing will reopen, allowing them to move north into central Europe. (Photo by Gregorio Borgia/AP Photo)

Habimana, 45, and her children walk to market to sell pottery on Idjwi island in the Democratic Republic of Congo, November 23, 2016. (Photo by Therese Di Campo/Reuters)