Coast and marine winner: Storm Gull (Lesser black-backed gull), New Haven, East Sussex. (Photo by Craig Denford/British Wildlife Photography Awards)

A frog appears to have a big smile for the camera, Russia. Things are heating up at the Comedy Wildlife Photography Awards as the shortlisted final 40 entries are revealed. This year's competition has featured over 2200 hilarious entries from around the world with photos including a fox face planting in the snow, a dancing brown bear and a pair of seemingly headless penguins – all beautifully photographed with perfect comedy timing and a strong conservation message. (Photo by Artyom Krivosheev/Barcroft Images/Comedy Wildlife Photography Awards 2016)

“Romantic”. A brown bear in Martinselkonen, Finland. (Photo by Valtteri Mulkahainen/Comedy Wildlife Photography Awards)

A golden huddle by Minqiang Lu, China. Two females and a male golden snub-nosed monkey huddle together to keep warm in the extreme cold. Threatened by forest loss and fragmentation, this endangered species is confined to central China. Restricted to living high up in the temperate forests, these monkeys – here in the Qinling mountains in Shaanxi province – feed mostly in the trees, on leaves, bark, buds and lichen. In heavy wind and snow, Minqiang walked up the mountain carrying his equipment. He stayed for half an hour in temperatures of –10C opposite the tree where the group was huddled before achieving this eye-level composition. (Photo by Minqiang Lu/Wildlife Photographer of the Year)

A zebra walks through the recently flooded Safari Park located at Safari World November 2, 2011 in Bangkok, Thailand. (Photo by Paula Bronstein /Getty Images)

Highly commended birds : Crested guan by Tim Hunt (UK). ‘This photo shows a crested guan in the cloud forests of Costa Rica as it pauses while foraging on a lone branch. Due to the clouds that are so typical for this habitat, I could often only photograph the bird’s silhouette against a grey sky. But then, for a short moment only, the sun broke through the clouds, and I overexposed the image by over two stops in order to blow out the background and allow this beautifully marked bird to stand out’. (Photo by Tim Hunt/GDT European Wildlife Photographer of the Year 2015)

Behaviour: Mammals category. Giant Gathering by Tony Wu, USA. Dozens of sperm whales mingled noisily off Sri Lanka’s northeast coast, stacked as far down as Tony could see. This was a congregation of dozens of social units, like a gathering of the clans. Aggregations like this could be a critical part of the whales’ rich social lives but are rarely reported. Some two thirds of the population was wiped out before commercial whaling was banned in 1986. This kind of major gathering could be “a sign that populations are recovering”, says Tony. (Photo by Tony Wu/Wildlife Photographer of the Year 2017)

Highly commended, mammals: Gelada after the storm – Marco Gaiotti (Italy). “Gelada baboons are the only monkey species in the world that feed on grasses. They are native to the tableland of Ethiopia. Every morning large family groups wander from their sleeping places in the steep rock face, up to 1,000 metres high, to the feeding grounds at the tablelands. This image clearly depicts their feeding strategy: they pull out bunches of grass, sort the stalks and then lift them to their mouth. This shot was taken towards the end of the rainy season after a heavy storm”. (Photo by Marco Gaiotti/2019 GDT European Wildlife Photographer of the Year)