Grand prize winner: environmental photographer of the year 2023; also 1st place in the humanity v nature category; and winner of the 2023 public award. Lopé national park, Gabon, 2021. An angry elephant tries to defend itself after it was hit by a train that crosses paths the animals use within Lopé national park. Park officials decided the elephant was too severely injured to be saved. After it was killed, the park director distributed the meat to local people. As the forest loses its carrying capacity to sustain its megafauna, this kind of human-wildlife conflict is increasing. (Photo by Jasper Doest/Environmental Photography Award)

A spotted owlet appears to be playing peek-a-boo near Rancharda Lake in Ahmedabad, India on February 9, 2016. The species is common across the Indian subcontinent. (Photo by Caters News Agency)

A polar bear fast asleep at London Zoo. (Photo by Fox Photos/Getty Images). 1961

An Indian Sikh Nihang, or warrior, performs Gatka martial arts skills during a procession at the Golden Temple in Amritsar on January 6, 2014, as part of birth anniversary celebrations of Guru Gobind Singh. The birth anniversary of Guru Gobind Singh, the tenth Sikh guru, is marked on January 6. (Photo by Narinder Nanu/AFP Photo)

Birds behaviour winner: Land of the Eagle by Audun Rikardsen, Norway. High on a ledge, on the coast near his home in northern Norway, Rikardsen carefully positioned an old tree branch that he hoped would make a perfect golden eagle lookout. To this, he bolted a tripod head with a camera, flashes and motion sensor attached, and built himself a hide a short distance away. From time to time, he left road‑kill carrion nearby. Very gradually – over the next three years – a golden eagle got used to the camera and started to use the branch regularly to survey the coast below. (Photo by Audun Rikardsen/2019 Wildlife Photographer of the Year)

Zhu Jianqiang, or “strong-willed pig”, famous for surviving more than a month buried in the ruins after the magnitude-8.0 Wenchuan earthquake in 2008, feeds on food sent by admirers wishing it a safe winter in Jianchuan Museum, Chengdu city, southwest China's Sichuan province, 11 January 2021. (Photo by Rex Features/Shutterstock)

Behaviour: Mammals category. Giant Gathering by Tony Wu, USA. Dozens of sperm whales mingled noisily off Sri Lanka’s northeast coast, stacked as far down as Tony could see. This was a congregation of dozens of social units, like a gathering of the clans. Aggregations like this could be a critical part of the whales’ rich social lives but are rarely reported. Some two thirds of the population was wiped out before commercial whaling was banned in 1986. This kind of major gathering could be “a sign that populations are recovering”, says Tony. (Photo by Tony Wu/Wildlife Photographer of the Year 2017)

Shooting star by Tony Wu, US/Japan. Winner, underwater. The electrifying reproductive dance of a giant sea star. As the surrounding water filled with sperm and eggs from spawning sea stars, Wu faced several challenges. Stuck in a small, enclosed bay with only a macro lens for photographing small subjects, he backed up to squeeze the undulating sea star into his field of view. The dancing posture of spawning sea stars rising and swaying may help release eggs and sperm, or may help sweep the eggs and sperm into the currents where they fertilise together in the water. Kinko Bay, Japan. (Photo by Tony Wu/Wildlife Photographer of the Year)