Commended. In late May, about a quarter of a million snow geese arrive from North America to nest on Wrangel Island, in northeastern Russia. They form the world's largest breeding colony of snow geese. Photographer Sergey Gorshkov spent two months on the remote island photographing the unfolding dramas. Arctic foxes take advantage of the abundance of eggs, caching surplus eggs for leaner times. But a goose (here the gander) is easily a match for a fox, which must rely on speed and guile to steal eggs. “The battles were fairly equal”, notes Sergey, “and I only saw a fox succeed in grabbing an egg on a couple of occasions, despite many attempts”. Surprisingly, “the geese lacked any sense of community spirit”, he adds, “and never reacted when a fox harassed a neighboring pair nesting close by”. (Photo by Sergey Gorshkov/Veolia Environnement Wildlife Photographer)

“Family disagreement”. (Photo by Vlado Pirsa/Comedy Wildlife Photo Awards 2019)

Hidden Britain category winner. Garden Spider by Alan Smith from Reading, Berkshire. (Photo by Alan Smith/British Wildlife Photography Awards/PA Wire Press Association)

A zebra walks through the recently flooded Safari Park located at Safari World November 2, 2011 in Bangkok, Thailand. (Photo by Paula Bronstein /Getty Images)

Behaviour: Mammals category. Giant Gathering by Tony Wu, USA. Dozens of sperm whales mingled noisily off Sri Lanka’s northeast coast, stacked as far down as Tony could see. This was a congregation of dozens of social units, like a gathering of the clans. Aggregations like this could be a critical part of the whales’ rich social lives but are rarely reported. Some two thirds of the population was wiped out before commercial whaling was banned in 1986. This kind of major gathering could be “a sign that populations are recovering”, says Tony. (Photo by Tony Wu/Wildlife Photographer of the Year 2017)

Highly commended, mammals: Gelada after the storm – Marco Gaiotti (Italy). “Gelada baboons are the only monkey species in the world that feed on grasses. They are native to the tableland of Ethiopia. Every morning large family groups wander from their sleeping places in the steep rock face, up to 1,000 metres high, to the feeding grounds at the tablelands. This image clearly depicts their feeding strategy: they pull out bunches of grass, sort the stalks and then lift them to their mouth. This shot was taken towards the end of the rainy season after a heavy storm”. (Photo by Marco Gaiotti/2019 GDT European Wildlife Photographer of the Year)

Highly commended birds : Crested guan by Tim Hunt (UK). ‘This photo shows a crested guan in the cloud forests of Costa Rica as it pauses while foraging on a lone branch. Due to the clouds that are so typical for this habitat, I could often only photograph the bird’s silhouette against a grey sky. But then, for a short moment only, the sun broke through the clouds, and I overexposed the image by over two stops in order to blow out the background and allow this beautifully marked bird to stand out’. (Photo by Tim Hunt/GDT European Wildlife Photographer of the Year 2015)

Nosy neighbour by Sam Hobson, UK. Sam knew exactly who to expect when he set his camera on the wall one summer’s evening in a suburban street in Bristol, the UK’s famous fox city. He wanted to capture the inquisitive nature of the urban red fox in a way that would pique the curiosity of its human neighbours about the wildlife around them. (Photo by Sam Hobson/2016 Wildlife Photographer of the Year)