Guardian of the Mangroves – Overall Winner. Tanya Houppermans, Cuba. A curious American crocodile (Crocodylus acutus) swims right up to Tanya, at Gardens of the Queen (Jardines De La Reina), an archipelago off the coast of Cuba. It has been strictly protected since 1996, and is one of the most untouched marine ecosystems in the world. ‍ “The healthy population of American crocodiles is down to the pristine condition of the mangroves and I wanted to capture close ups of this gentle giant in its natural habitat. I hope this image can illustrate that protecting areas like this is so critical”. (Photo by Tanya Griffin Houppermans/Mangrove Photographer of the Year)

Long-tailed mayflies (Palingenia longicauda) at Tisza river near Tiszainoka 135km (84 miles) southeast of Budapest, June 23, 2013. Millions of these short-lived mayflies engage in a frantic rush to mate and reproduce before they perish in just a few hours during “Tiszaviragzas” or Tisza blooming season from late spring to early summer every year. (Photo by Laszlo Balogh/Reuters)

A baby rhesus macaque (Macaca mulatta) looks up as it plays with a tiger cub at a zoo in Hefei, Anhui province, August 2, 2012. (Photo by Jianan Yu/Reuters)

The Enemy, 1944. With the outbreak of world war two, photojournalism was enlisted in defence of Mother Russia. In this photo by Anatoli Egorov, who was wounded in action, corporal Stepan Vasiljevich Ovcharenko shoots at enemy troops with a machine gun. (Photo by Anatoli Egorov/Lumiere Brothers Center for Photography)

Deputies of the opposition fight with deputies of the majority for a second time in two days, during the second session of the newly elected Ukrainian parliament in Kiev on December 13, 2012. (Photo by Sergei Supinsky/AFP Photo)

People smeared with “Gulal” or coloured powder during celebrations of Hindu spring festival “Holi” in Hyderabad, India on March 7, 2023. (Photo by Noah Seelam/AFP Phoot)

A competition, now in its 43rd year, dedicated to showcasing the beautiful and bizarre as seen under a light microscope attracted over 2,000 entries from 88 countries. Here: Honorable Mention by Emre Can Alagöz, Istanbul, Turkey: The eyes of a jumping spider, magnified 6x. (Photo by Emre Can Alagöz/2017 Nikon Small World Photomicrography Competition)

Respect, Kronotsky nature reserve, Russia. The photographer’s cat, Ryska – her name means little lynx in Russian – stands outside their cabin and with aggressive posturing warns off a fox. In winter, foxes would regularly visit the cabin searching for food. If one peered in at the window, possible when the snow was deep, Ryska would sit on the other side, fur raised, and growl. When outside, she would hold her ground. The foxes were not always frightened and so encounters could be a sort of dance. (Photo by Igor Shpilenok/Unforgettable Behaviour/NHM)