Highly commended birds : Crested guan by Tim Hunt (UK). ‘This photo shows a crested guan in the cloud forests of Costa Rica as it pauses while foraging on a lone branch. Due to the clouds that are so typical for this habitat, I could often only photograph the bird’s silhouette against a grey sky. But then, for a short moment only, the sun broke through the clouds, and I overexposed the image by over two stops in order to blow out the background and allow this beautifully marked bird to stand out’. (Photo by Tim Hunt/GDT European Wildlife Photographer of the Year 2015)

Underwater photographer of the year – winner. Dancing Octopus by Gabriel Barathieu (France). Location: Island of Mayotte, off the coast of south-east Africa. “Balletic and malevolent”, one judge said of this octopus, hunting in a lagoon. Barathieu waited until spring tides when there was just 30cm of water on the flats and plenty of light in the shallows. (Photo by Gabriel Barathieu/UPY2017)

A hyena kisses his own reflection while appearing to walk on water. (Photo by Jake De Wet/Caters News Agency)

Shortlisted: “Two big eyes” by Miao Yong (Zejiang province, China). Damselflies look over the leaves. “I was photographing insects in a park near my home when suddenly I found two damselflies in the grass. They kept flying and it was very difficult to focus until suddenly they parked behind a leaf”. (Photo by Miao Yong/2017 Royal Society of Biology Photographer of the Year)

A leg of a “diablito” frog (Oophaga sylvatica) is photographed in a laboratory at a laboratory in the zoo of Cali, Colombia, on July 19, 2019. Colombia is the second country with the largest number of amphibians in the world after Brazil. More than 40% of amphibian species worldwide are in danger of extinction. (Photo by Luis Robayo/AFP Photo)

A red squirrel and a woodpecker having an “argument” over some nuts in a woodland near Lockerbie, Dumfries and Galloway in Johnsfield, England in September 2020. (Photo by Karen Crawford/South West News Service)

This is the mesmerising image of a colourful Milky Way shining down on Mount Fuji. Steve Lance Lee, 37, captured the sky above the iconic mountain whilst on his honey moon in Lake Kawaguchiko, Japan. Mount Fuji is illuminated underneath the galaxy of stars, creating an impressive landscape. Here: The beautiful image captured by Steve Lance Lee of the Milky Way seemingly errupting from Mt. Fuji in Japan. (Photo by Steve Lance Lee/Caters News)

The males, of the Oriental Garden species, are battling over territory in Solapur in the Maharashtra state, India on May 20, 2022. Occasionally, the tussle can be brutal, ending in a fatality with one lizard biting another to encourage them to retreat. (Photo by Ratnakar Hiremath/Solent News)