A golden huddle by Minqiang Lu, China. Two females and a male golden snub-nosed monkey huddle together to keep warm in the extreme cold. Threatened by forest loss and fragmentation, this endangered species is confined to central China. Restricted to living high up in the temperate forests, these monkeys – here in the Qinling mountains in Shaanxi province – feed mostly in the trees, on leaves, bark, buds and lichen. In heavy wind and snow, Minqiang walked up the mountain carrying his equipment. He stayed for half an hour in temperatures of –10C opposite the tree where the group was huddled before achieving this eye-level composition. (Photo by Minqiang Lu/Wildlife Photographer of the Year)

A huge atlas moth, with a wingspan of more than 9in, photographed on an areca nut plantation in Sirsi, India. (Photo by Uday Hegde/Close Up Photographer of the Year)

Grand title winner: Environmental photographer of the year. The Bitter Death Of Birds by Mehdi Mohebi Pour. This photo shows the efforts of the environmental forces to collect the bodies and prevent the spread of this disease. The Miankaleh wetland is being destroyed by changes in the climate and it is my duty as a photographer to highlight these problems and create a record for history. I want to prevent the complete destruction of the wetland and the potential environmental disaster by showing the issues and threats to these beautiful natural places. (Photo by Mehdi Mohebi Pour/Environmental Photographer of the Year)

Guardian of the Mangroves – Overall Winner. Tanya Houppermans, Cuba. A curious American crocodile (Crocodylus acutus) swims right up to Tanya, at Gardens of the Queen (Jardines De La Reina), an archipelago off the coast of Cuba. It has been strictly protected since 1996, and is one of the most untouched marine ecosystems in the world. ‍ “The healthy population of American crocodiles is down to the pristine condition of the mangroves and I wanted to capture close ups of this gentle giant in its natural habitat. I hope this image can illustrate that protecting areas like this is so critical”. (Photo by Tanya Griffin Houppermans/Mangrove Photographer of the Year)

“Romantic”. A brown bear in Martinselkonen, Finland. (Photo by Valtteri Mulkahainen/Comedy Wildlife Photography Awards)

Shooting star by Tony Wu, US/Japan. Winner, underwater. The electrifying reproductive dance of a giant sea star. As the surrounding water filled with sperm and eggs from spawning sea stars, Wu faced several challenges. Stuck in a small, enclosed bay with only a macro lens for photographing small subjects, he backed up to squeeze the undulating sea star into his field of view. The dancing posture of spawning sea stars rising and swaying may help release eggs and sperm, or may help sweep the eggs and sperm into the currents where they fertilise together in the water. Kinko Bay, Japan. (Photo by Tony Wu/Wildlife Photographer of the Year)

Human Connection Award: People and Planet Ocean – Winner – Steve Woods. A free diver interacts with a sperm whale among a cloud of sargassum weed, Dominica. (Photo by Steve Woods/Ocean Photographer of the Year 2022)

Portrait category, bronze award winner. Purple-crested Turaco Gallirex porphyreolophus. Lower Mpushini, near Pietermaritzburg, KwaZulu-Natal, South Africa. (Photo by Richard Flack/Bird Photographer of the Year 2022)