Behaviour winner; Courting Royals: two royal terns in courtship display by Kristian Bell. Another beautiful morning on a beautiful beach on the Gulf Coast of Florida seemed to prompt these two royal terns to commence an intricate courtship dance. The photograph was taken with a Canon 300mm lens and 2x extender. (Photo by Kristian Bell/Deakin University/Royal Society Publishing Photography Competition 2018)

Honourable mention, Behaviour category. Toss the scorpion – Indian roller playing with its kill by Susmita Datta. The image was taken during an early morning safari drive at Tadoba Andhari Tiger Reserve in India. (Photo by Susmita Datta/PA Wire/Royal Society Publishing Photography Competition 2017)

Runner-up. “The heavens opened and the streets around Shibuya, Tokyo, were suddenly drenched in even more colour and movement. I was taking photos (my friend was holding the umbrella) when I noticed this woman at the crossing. I’m pleased with the futuristic feel of the image”. (Photo by Katherine Bridgestock/The Guardian)

The photography duo of Floto+Warner created the series, “Colorant”, from an idea that stemmed out of a previous series and the fascination of landscapes, with results that leave one in awe. Creating shapes, not experienced in nature, they tossed colored water in the air to capture “a momentary graffiti of air and space”. Using a high shutter speed to capture these fleeting moments, Floto/Warner has produced a multi-medium series with jaw-dropping results. Photo: “Orange Salt Flats”. (Photo by Floto/Warner)