A frog appears to have a big smile for the camera, Russia. Things are heating up at the Comedy Wildlife Photography Awards as the shortlisted final 40 entries are revealed. This year's competition has featured over 2200 hilarious entries from around the world with photos including a fox face planting in the snow, a dancing brown bear and a pair of seemingly headless penguins – all beautifully photographed with perfect comedy timing and a strong conservation message. (Photo by Artyom Krivosheev/Barcroft Images/Comedy Wildlife Photography Awards 2016)

Japanese macaque monkeys enjoy sitting in the hot springs at Jigokudani-Onsen (Hell Valley) on January 23, 2005 in Jigokudani, Nagano-Prefecture, Japan. Japanese Macaques, also known as snow monkeys are the most northerly nonhuman primate in the world. In 1963 a female Macaque ventured into the hot springs to retrieve some soybeans. This behaviour was adopted by other monkeys, and eventually by the entire troop. This Macaque troop regularly visits the Jigokudani-Onsen springs to escape the cold. The hot springs are said to help relieve nerve pain and fatigue. (Photo by Koichi Kamoshida/Getty Images)

A combination picture shows Will Adam, 14, snowboarding down a street on Beacon Hill during a large winter blizzard in Boston, Massachusetts, United States January 27, 2015 (top), and a woman looking at her phone as she walks down the same street June 13, 2015. Outdoor scenes in Boston show local residents enjoying balmy weather amid lush greenery as the summer solstice approaches on 21 June. The very same locations suffered heavy snowstorms last winter, with snow ploughs, skiers and snowboarders battling the drifts. Boston got 275.8 cm of snow over the winter, the most since 1872, when records began. A few months after the snowstorms, Brian Snyder revisited the same places and shot pictures at exactly the same locations. (Photo by Brian Snyder/Reuters)

A tale of two foxes. Surprising behaviour, witnessed in Wapusk national park, on Hudson Bay, Canada, in early winter. Red foxes don’t actively hunt Arctic foxes, but where the ranges of two predators overlap, there can be conflict. Though the light was poor, the snow-covered tundra provided the backdrop for the moment that the red fox paused with the smaller fox in its mouth in a grim pose. (Photo by Don Gutoski/2015 Wildlife Photographer of the Year)

A newly wed couple jumps to pose for their wedding photographs after their group wedding ceremony which was held as part of the Harbin International Ice and Snow Festival in the northern city of Harbin, Heilongjiang province January 6, 2015. The wedding ceremony was organised by the city government and 11 couples from China, Russia and Egypt participated in the ceremony. (Photo by Kim Kyung-Hoon/Reuters)

Tourists photograph a red dear stag visiting a car park near Glen Coe on December 1, 2017 in Glen Coe,Scotland. On the first day of the meteorological calendar, the UK was experiencing slightly warmer temperatures today with weather forecasters indicating that the recent cold spell is almost over. Yesterday's snow across eastern parts of England is expected to melt away over the course Friday as temperatures start to rise. (Photo by Jeff J. Mitchell/Getty Images)

Respect, Kronotsky nature reserve, Russia. The photographer’s cat, Ryska – her name means little lynx in Russian – stands outside their cabin and with aggressive posturing warns off a fox. In winter, foxes would regularly visit the cabin searching for food. If one peered in at the window, possible when the snow was deep, Ryska would sit on the other side, fur raised, and growl. When outside, she would hold her ground. The foxes were not always frightened and so encounters could be a sort of dance. (Photo by Igor Shpilenok/Unforgettable Behaviour/NHM)