Nature – first prize, stories. Pandemic Pigeons – A Love Story. The photographer’s daughter, Merel, cowers after Dollie flies past and perches on the balcony before entering the house in Vlaardingen in the Netherlands on 6 April 2020. “She’s still frightened when Dollie suddenly lands on the balcony railing. I hide my smile behind the camera, as I try to comfort her by saying they won’t hurt you. “I thought he was going to attack me”, she replies. As the nesting pigeons keep coming back to our place, slowly my girls have started to appreciate them – perhaps not as much as I do, but it’s a start”. (Photo by Jasper Doest/World Press Photo 2021)

With its huge eyes, comical name and diminutive size, Mark R. Smith’s image of a baby Hawaiian bobtail squid can’t help but raise a smile. A curiously endearing creature, the cephalopod is just 1.5cm across, its mantle cavity bearing more than a passing resemblance to a rather natty shower cap. But it is also a beautiful example of symbiosis – nature’s version of “I’ll scratch your back if you scratch mine” – for on the underside of the squid is a light organ which houses bioluminescent bacteria. The squid offers the bacteria protection and food, while the bacteria emit a glow – a handy trait that the squid uses to offset its silhouette, helping it to evade predators in the depths below. Mark R. Smith’s entry combines several images of a Hawaiian bobtail squid with different focus lengths to create a final picture with greater depth of field than normal. (Photo by Mark R. Smith/Wellcome Images/Macroscopic Solutions)