Shooting star by Tony Wu, US/Japan. Winner, underwater. The electrifying reproductive dance of a giant sea star. As the surrounding water filled with sperm and eggs from spawning sea stars, Wu faced several challenges. Stuck in a small, enclosed bay with only a macro lens for photographing small subjects, he backed up to squeeze the undulating sea star into his field of view. The dancing posture of spawning sea stars rising and swaying may help release eggs and sperm, or may help sweep the eggs and sperm into the currents where they fertilise together in the water. Kinko Bay, Japan. (Photo by Tony Wu/Wildlife Photographer of the Year)

A tale of two foxes. Surprising behaviour, witnessed in Wapusk national park, on Hudson Bay, Canada, in early winter. Red foxes don’t actively hunt Arctic foxes, but where the ranges of two predators overlap, there can be conflict. Though the light was poor, the snow-covered tundra provided the backdrop for the moment that the red fox paused with the smaller fox in its mouth in a grim pose. (Photo by Don Gutoski/2015 Wildlife Photographer of the Year)

Arctic treasure by Sergey Gorshkov (Russia). An arctic fox carries its egg trophy from a raid on a snow goose nest and heads for a suitable burial spot. Finalist 2017, Animal Portraits. (Photo by Sergey Gorshkov/2017 Wildlife Photographer of the Year)

Army paratroopers carry the casket of Private Howard at the Wellington Cathedral on December 21, 2010 in Wellington, New Zealand. Private Howard was serving with the British army when was killed by friendly fire from a US Air Force plane in Helmand Province, Afghanistan earlier this month. Private Howard is the fifth NZ-born soldier to be killed in action in Afghanistan. (Photo by Marty Melville/Getty Images)

A protester holds a poster displaying the message “Get away UK” during an anti-British demonstration in the Iranian capital that resulted in a break in at the British Embassy on November 29, 2011 in Tehran, Iran. (Photo by FarsNews/Getty Images)

Actress Olga Kurylenko poses for a photograph at the British Academy of Film and Arts (BAFTA) awards ceremony at the Royal Opera House in London February 16, 2014. (Photo by Suzanne Plunkett/Reuters)

I had to stay late at work, Chubut, Argentina. “South sea elephant in Patagonia (Isla Escondida) They adopt very curious gestures!”. (Photo by Luis Burgue/Comedy Wildlife Photography Awards 2020)

Winner – Animal Portraits: The pose by Mogens Trolle, Denmark. A young male proboscis monkey cocks his head slightly and closes his eyes. Unexpected pale blue eyelids now complement his immaculately groomed auburn hair. He poses for a few seconds as if in meditation. He is a wild visitor to the feeding station at Labuk Bay Proboscis Monkey Sanctuary in Sabah, Borneo – “the most laid-back character”, says Trolle, “quite unlike anything I’ve ever seen on another monkey” – connects us, he hopes, with a fellow primate. (Photo by Mogens Trolle/Wildlife Photographer of the Year 2020)