Shortlisted: Pooyan Shadpoor, Houcheraghi. While walking along the shore of Larak, Iran – an island in the Persian Gulf – Shadpoor came across this luminous scene. The “magical lights of (the) plankton ... enchanted me so that I snapped the shot”, he writes. (Photo by Pooyan Shadpoor/2016 EPOTY)

Second place, Nature: double trapping. Picture taken in the Brazilian Pantanal ... when I downloaded the CF did not want to believe it ... The nature knows we always give magnificent events but sometimes extraordinary. (Photo by Massimiliano Bencivenni/National Geographic Travel Photographer of the Year Contest)

Breath of an Arctic fox by Marco Gaiotti, Italy. Marco was watching this little Arctic fox as it incessantly called another nearby. Gradually he noticed the fox’s wet breath was quickly freezing in the air after each call. It was late winter in Spitsbergen, Svalbard, and the air was -35C (-31F). Photographing Arctic foxes is often frustrating, as they are normally running around fast in search of food, but this one was very relaxed and let Marco get close enough to focus on it, with the light glowing perfectly in the background. (Photo by Marco Gaiotti/Wildlife Photographer of the Year 2021)

Aurorae category runner-up: Lone Tree under a Scandinavian Aurora by Tom Archer (UK). The photographer decided to explore the area around the hotel on a very crisp -35C evening in Finnish Lapland. When he found this tree, he decided to wait for the misty conditions to change and could not believe his luck when the sky cleared and the aurora came out in the perfect spot. Archer spent about an hour photographing it before his camera started to lock up because of the harsh conditions, but by then he was happy to call it a night. (Photo by Tom Archer/2020 Astronomy Photographer of the Year)

M42 Subtle V1 cropped. One of the most well-known astronomical objects in our universe is the Orion Nebula and this image depicts the wider region of the Orion Molecular Cloud Complex that is its home. This complex also includes another popular target for astrophotographers, the Horsehead Nebula, as well as Barnard’s Loop and the Running Man Nebula, which can be seen to the left of this photograph. (Photo by Patrick Gilliland)

Highly commended birds : Crested guan by Tim Hunt (UK). ‘This photo shows a crested guan in the cloud forests of Costa Rica as it pauses while foraging on a lone branch. Due to the clouds that are so typical for this habitat, I could often only photograph the bird’s silhouette against a grey sky. But then, for a short moment only, the sun broke through the clouds, and I overexposed the image by over two stops in order to blow out the background and allow this beautifully marked bird to stand out’. (Photo by Tim Hunt/GDT European Wildlife Photographer of the Year 2015)

A) 1st place WINNER – Roy Rimmer. “This rat was in an outdoor set I made, the set up is two meters long and a meter wide made of Perspex,it has a plywood front with holes cut in for my camera and flash guns, I placed two rusty paint cans in the set up and the rat would leap from one can too the other, I had to use flash to freeze the action”.

Behaviour: Mammals category. Giant Gathering by Tony Wu, USA. Dozens of sperm whales mingled noisily off Sri Lanka’s northeast coast, stacked as far down as Tony could see. This was a congregation of dozens of social units, like a gathering of the clans. Aggregations like this could be a critical part of the whales’ rich social lives but are rarely reported. Some two thirds of the population was wiped out before commercial whaling was banned in 1986. This kind of major gathering could be “a sign that populations are recovering”, says Tony. (Photo by Tony Wu/Wildlife Photographer of the Year 2017)