Highly commended birds : Crested guan by Tim Hunt (UK). ‘This photo shows a crested guan in the cloud forests of Costa Rica as it pauses while foraging on a lone branch. Due to the clouds that are so typical for this habitat, I could often only photograph the bird’s silhouette against a grey sky. But then, for a short moment only, the sun broke through the clouds, and I overexposed the image by over two stops in order to blow out the background and allow this beautifully marked bird to stand out’. (Photo by Tim Hunt/GDT European Wildlife Photographer of the Year 2015)

Enjoying a tickle from her keeper, an orphaned orangutan at the SOCP Quarantine Centre in Sumatra, Indonesia on March 20, 2016. The island’s orangutan population has been devastated in recent years as the forest areas are destroyed to make way for palm oil plantations. (Photo by Jami Tarris/Barcroft Media)

Shortlisted. Dragonfly, North York Moors national park, by Jonathan Green: “In June I was at May Beck with some colleagues when someone spotted this Hawker perched in a gorse bush. I love the detail on the wings, and it’s rare to be able to get so much of such a small subject in clear focus. Getting a few scratches was worth it”. (Photo by Jonathan Green/2020 UK National Parks Photography Competition)

A tale of two foxes. Surprising behaviour, witnessed in Wapusk national park, on Hudson Bay, Canada, in early winter. Red foxes don’t actively hunt Arctic foxes, but where the ranges of two predators overlap, there can be conflict. Though the light was poor, the snow-covered tundra provided the backdrop for the moment that the red fox paused with the smaller fox in its mouth in a grim pose. (Photo by Don Gutoski/2015 Wildlife Photographer of the Year)

It’s enough to make your head spin. The photographer Katherine Young set out to shoot spiral staircases in London, England to great effect, including this shot she calls the Downward Spiral Part III. (Photo by Katherine Young/Rex Features/Shutterstock)

As an American in 1990s Cuba, Tria Giovan risked being branded a traitor. But the photographer continued to visit and, from the dance hall to the hair salon, she captured the resilient spirit of the Cuban people. The 120 images in Tria Giovan’s “The Cuba Archive” are from the period in the 90s when, as an American, travel to Cuba could have seen her branded a traitor, as the country was subject to a US trade embargo. Her trip required lots of planning – and patience. (Photo by Tria Giovan/The Guardian)

Tribal artists and supporters from the southern Indian state of Karnataka dance outside the residence of Congress Party President Sonia Gandhi during her birthday celebrations in New Delhi, India, Wednesday, December 9, 2015. The Italian-born Gandhi is the widow of former Prime Minister Rajiv Gandhi and has led the Congress party since 1998. (Photo by Altaf Qadri/AP Photo)

Two women lighting cigarettes on a tennis court in Essex, England circa 1930's. (Photo by Keystone View/FPG/Getty Images)