“Family disagreement”. (Photo by Vlado Pirsa/Comedy Wildlife Photo Awards 2019)

Hidden Britain category winner. Garden Spider by Alan Smith from Reading, Berkshire. (Photo by Alan Smith/British Wildlife Photography Awards/PA Wire Press Association)

Professional conceptual category winner. Greetings from Mars, by Julien Mauve, France. Mauve says: “I have always wondered what it would be like to discover a totally different world ... and to photograph it for the first time as if I was Ansel Adams. So I came up with this project, which is about space exploration and discovery. But it’s also about our behavior in front of landscapes and how we create pictures that will share our personal story with the world”. (Photo by Julien Mauve)

Runner-up. “The City of London, looking towards the Royal Exchange and the Bank of England. The ever-changing London skyline provides many excellent opportunities for cityscape photography, none more exciting than the ebb and flow of traffic at night”. (Photo by Mark Caldon/The Guardian)

Novice DSLR (Digital single-lens reflex camera), 2nd Place. “Smile of a Friend”, American Crocodile (Crocodylus acutus) in Jucaro, Cuba. (Photo by Antonio Pastrana/The Ocean Art 2018 Underwater Photography Competition)

Runner-up; Hon Yen marine ecosystem. Phu Yen, Vietnam. Every year between May and August, the coral of this rich and diverse ecosystem becomes exposed at low tide. (Photo by Truong Hoai Vu/Royal Society of Biology Photography Competition)

October winner. “A burst of the aurora borealis over Kirkjufell, taken in September 2016. This shot has been a holy grail for me and was finally captured after many trips”. MICK RYAN, JUDGE: Iceland is a hotspot for aurora photography and this composition of the symmetric, free-standing Kirkjufell and the trident waterfall is much-sought after. If you can time your visit when the aurora “kp” index is high and know the settings for nighttime aurora images, you may end up with a beautiful photograph like this. (Photo by DВ/The Guardian)

This year’s overall winner and winner of the coast and marine category is George Stoyle with his image “Hitchhikers” of a Lion’s mane jellyfish, photographed at St Kilda, off the Island of Hirta, Scotland. (Photo by George Stoyle/British Wildlife Photography Awards 2016)