First place, Portrait. Three flat-coated retrievers – Crew, Darcie and Pagan – by Carol Durrant from the UK. (Photo by Carol Durrant/Dog Photographer of the Year 2018)

Aurora Australis from Beerbarrel Beach, by James Stone. Runner Up: Aurorae. (Photo by James Stone/Astronomy Photographer of the Year)

Birds behaviour winner: Land of the Eagle by Audun Rikardsen, Norway. High on a ledge, on the coast near his home in northern Norway, Rikardsen carefully positioned an old tree branch that he hoped would make a perfect golden eagle lookout. To this, he bolted a tripod head with a camera, flashes and motion sensor attached, and built himself a hide a short distance away. From time to time, he left road‑kill carrion nearby. Very gradually – over the next three years – a golden eagle got used to the camera and started to use the branch regularly to survey the coast below. (Photo by Audun Rikardsen/2019 Wildlife Photographer of the Year)

“Photographer Seth Casteel has definitely taken animal photography to new heights – or depths – with his hilarious photos of dogs chasing balls underwater. Casteel, who, naturally, has two dogs of his own (a mini Labradoodle and a Norwich terrier), said that he's drawn to underwater dog photography because the photos show a side of dogs that people may have never seen before”.

Photo: Seth Casteel Underwater Dog Pictures (Photo by Seth Casteel)

Shooting star by Tony Wu, US/Japan. Winner, underwater. The electrifying reproductive dance of a giant sea star. As the surrounding water filled with sperm and eggs from spawning sea stars, Wu faced several challenges. Stuck in a small, enclosed bay with only a macro lens for photographing small subjects, he backed up to squeeze the undulating sea star into his field of view. The dancing posture of spawning sea stars rising and swaying may help release eggs and sperm, or may help sweep the eggs and sperm into the currents where they fertilise together in the water. Kinko Bay, Japan. (Photo by Tony Wu/Wildlife Photographer of the Year)

A huge atlas moth, with a wingspan of more than 9in, photographed on an areca nut plantation in Sirsi, India. (Photo by Uday Hegde/Close Up Photographer of the Year)

Komodo Judo by Andrey Gudkov. Finalist, Amphibians & Reptiles. Two large male Komodo dragons hissing angrily at each other in Indonesia’s Komodo national park. Komodo dragons can grow up to 8ft. (Photo by Andrey Gudkov/Wildlife Photographer of the Year 2015)

A pool ingeniously filled with clear water in the middle of Lake Kerniki in Greece enabled this shot of feeding pelicans. (Photo by Bence Mate/Close Up Photographer of the Year 2020)