Runner-up; Hon Yen marine ecosystem. Phu Yen, Vietnam. Every year between May and August, the coral of this rich and diverse ecosystem becomes exposed at low tide. (Photo by Truong Hoai Vu/Royal Society of Biology Photography Competition)

Birds behaviour winner: Land of the Eagle by Audun Rikardsen, Norway. High on a ledge, on the coast near his home in northern Norway, Rikardsen carefully positioned an old tree branch that he hoped would make a perfect golden eagle lookout. To this, he bolted a tripod head with a camera, flashes and motion sensor attached, and built himself a hide a short distance away. From time to time, he left road‑kill carrion nearby. Very gradually – over the next three years – a golden eagle got used to the camera and started to use the branch regularly to survey the coast below. (Photo by Audun Rikardsen/2019 Wildlife Photographer of the Year)