“Entwined Lives”. Tim Laman, US Winner, Wildlife photographer of the year. A young male orangutan makes the 30-metre climb up the thickest root of the strangler fig high above the canopy in Gunung Palung national park, one of the few protected orangutan strongholds in Indonesian Borneo. Laman had to do three days of climbing to position several GoPro cameras that he could trigger remotely. This shot was the one he had long visualised, looking down on the orangutan within its forest home. (Photo by Tim Laman/2016 Wildlife Photographer of the Year)

The National Geographic Photo Ark is a travelling exhibition of photographer Joel Sartore’s quest to create a photo archive of biodiversity around the world. So far, Sartore has captured studio portraits of more than 6,000 species – a number that he hopes to double. On 1 July, the ark will open at Melbourne zoo – the first time it has been exhibited in the southern hemisphere. More than 50 portraits will be on display, including many of Australian endangered animals being protected by programs at the zoo itself. These captions have been edited from text supplied by Melbourne zoo. Here: Barking owl. (Photo by Joel Sartore/National Geographic Photo Ark/The Guardian)

A lone male cheetah is set upon by a pack of African wild dogs. Peter Haygarth had been following the dogs as they hunted in Zimanga Private Game Reserve in KwaZulu-Natal, South Africa. On first encountering the cheetah, the dogs were wary, but as the rest of the pack arrived, their confidence grew and they began to encircle the cat. Peter kept his focus on the cat’s face. In a few minutes the spat was over as the cheetah fled. (Behaviour: mammals category). (Photo by Peter Haygarth)

“Eye of a toad”. Animal Portraits, Łukasz Bożycki, Poland. Early spring sees a pond near Łukasz’s home city of Warsaw, Poland, full of mating frogs and a few toads. On this March day, Łukasz shared the pond with them for an evening, sitting in the icy water in his chest-high waders, keeping as still as possible, despite the numbing cold, so that the amphibians could get used to him. “I wanted to find a fresh way of portraying the amphibians”, he says, “at water level”. Using a telephoto lens, he focused on one lone toad and waited for the sun to dip almost below the horizon before pressing the shutter, using flash to bring out the details in the shadow. His prize was “the glorious pool of sunset colour” and fiery glow of the toad’s eye. Nikon D80 + 70-300mm f4.5-5.6 lens + extension tube; 1/125 sec at f9 (-2.3 e/v); ISO 100; built-in flash. (Photo by Łukasz Bożycki)

Tim Laman is a field biologist and wildlife photojournalist. His pioneering research in the rain forest canopy in Borneo led to a PhD from Harvard and his first National Geographic article in 1997. Since then, he has pursued his passion for exploring wild places and documenting little-known and endangered wildlife by becoming a regular contributor to National Geographic. He has eighteen articles to his credit to date, all of which have had a conservation message. Some have focused on endangered species such as Orangutans or Hornbills, while others, such as a series of articles on Conservation International’s Biodiversity Hotspots, have highlighted regions under intense pressure.

A Puffin returns to it's nest with a beak full of sand eels on June 25, 2011 on Inner Farne, England. The Farne Islands, which are run by the National Trust, are situated two to three miles off the Northumberland coastline. The archipeligo of 16-28 separate islands (depending on the tide) make the summer home to approximately 100,000 pairs of breeding seabirds including around 36,000 Puffins, 32,000 Guillemots and 2,000 pairs of Arctic Terns. The species of birds which nest in internationally important numbers include Shag, Sandwich Tern and Arctic Tern. The coastline around The Farnes are also the breeding ground to one of Europe's largest Grey Seal colonies with around 4,000 adults giving birth to 1500 pups every year. (Photo by Dan Kitwood/Getty Images)

More than 1,500 snappers submitted their most hilarious pictures of all creatures great and small, and now 45 have made the cut. From drunken-eyed owls to embarrassed chipmunks and laughing goats – the finalists in the Comedy Wildlife Photography Awards are guaranteed to raise a smile. Here: 'Dancing sifaka'. (Photo by Alison Buttigieg/Comedy Wildlife Photography Awards/Mercury Press)

Polar frame, by Dmitry Kokh, Russia. When Kokh’s boat approached the small island of Kolyuchin in the Russian High Arctic, which had been abandoned by humans since 1992, he was surprised to spot movement in one of the houses. Binoculars revealed polar bears – more than 20 in total – exploring the ghost town. Dmitry used a low-noise drone to document them. (Photo by Dmitry Kokh/Wildlife Photographer of the Year 2022)