Shortlisted: Pooyan Shadpoor, Houcheraghi. While walking along the shore of Larak, Iran – an island in the Persian Gulf – Shadpoor came across this luminous scene. The “magical lights of (the) plankton ... enchanted me so that I snapped the shot”, he writes. (Photo by Pooyan Shadpoor/2016 EPOTY)

Second place, Nature: double trapping. Picture taken in the Brazilian Pantanal ... when I downloaded the CF did not want to believe it ... The nature knows we always give magnificent events but sometimes extraordinary. (Photo by Massimiliano Bencivenni/National Geographic Travel Photographer of the Year Contest)

Nosy neighbour by Sam Hobson, UK. Sam knew exactly who to expect when he set his camera on the wall one summer’s evening in a suburban street in Bristol, the UK’s famous fox city. He wanted to capture the inquisitive nature of the urban red fox in a way that would pique the curiosity of its human neighbours about the wildlife around them. (Photo by Sam Hobson/2016 Wildlife Photographer of the Year)

The Mount Bromo volcano erupts in the Bryce Canyon, Utah. (Photo by Reynold Dewantara/2016 National Geographic Travel Photographer of the Year Contest)

Under Exposed: Pekka Tuuri (Finland) – Great white shark, Isla Guadalupe, Mexico. “Isla Guadalupe is the world capital when it comes to observing great white sharks, but cage diving seriously limits the possibilities to take ‘fresh’ pictures. When I took this, the water close to the surface was quite milky, making photography very challenging. From out of the “mist”, I saw this great white shark lurking behind a school of fusiliers. I quickly focused on the shark and set a wide aperture to get focus blur on the fish, along with a fast shutter to avoid excessive motion blur”. (Photo by Pekka Tuuri/The Outdoor Photographer of the Year/The Guardian)

A lone male cheetah is set upon by a pack of African wild dogs. Peter Haygarth had been following the dogs as they hunted in Zimanga Private Game Reserve in KwaZulu-Natal, South Africa. On first encountering the cheetah, the dogs were wary, but as the rest of the pack arrived, their confidence grew and they began to encircle the cat. Peter kept his focus on the cat’s face. In a few minutes the spat was over as the cheetah fled. (Behaviour: mammals category). (Photo by Peter Haygarth)

Creative imagery bronze winner. Make-up by Chengbo Sun, China. (Photo by BPOTY/Cover Images)

Respect, Kronotsky nature reserve, Russia. The photographer’s cat, Ryska – her name means little lynx in Russian – stands outside their cabin and with aggressive posturing warns off a fox. In winter, foxes would regularly visit the cabin searching for food. If one peered in at the window, possible when the snow was deep, Ryska would sit on the other side, fur raised, and growl. When outside, she would hold her ground. The foxes were not always frightened and so encounters could be a sort of dance. (Photo by Igor Shpilenok/Unforgettable Behaviour/NHM)