A lone male cheetah is set upon by a pack of African wild dogs. Peter Haygarth had been following the dogs as they hunted in Zimanga Private Game Reserve in KwaZulu-Natal, South Africa. On first encountering the cheetah, the dogs were wary, but as the rest of the pack arrived, their confidence grew and they began to encircle the cat. Peter kept his focus on the cat’s face. In a few minutes the spat was over as the cheetah fled. (Behaviour: mammals category). (Photo by Peter Haygarth)

“Entwined Lives”. Tim Laman, US Winner, Wildlife photographer of the year. A young male orangutan makes the 30-metre climb up the thickest root of the strangler fig high above the canopy in Gunung Palung national park, one of the few protected orangutan strongholds in Indonesian Borneo. Laman had to do three days of climbing to position several GoPro cameras that he could trigger remotely. This shot was the one he had long visualised, looking down on the orangutan within its forest home. (Photo by Tim Laman/2016 Wildlife Photographer of the Year)

Winner, photojournalism. Elephant in the room, by Adam Oswell, Australia Zoo. Visitors watch a young elephant performing underwater. Oswell was disturbed by this scene, and organisations concerned with the welfare of captive elephants say performances like this encourage unnatural behaviour. In Thailand, there are now more elephants in captivity than in the wild. With the Covid pandemic causing tourism to collapse, elephant sanctuaries are becoming overwhelmed with animals that can no longer be looked after by their owners. (Photo by Adam Oswell/Wildlife Photographer of the Year 2021)

An innovative photographer attached a camera to a remote-controlled car, allowing him to capture angles of wild lions, rhinos and other animals. Over the last 11 years, Chris Bray has been taking pictures of animals using his toy car contraption while he takes guests on photography tours in Kenya. (Photo by Chris Bray/Caters News Agency)

An arctic tern feeds its chick on the Farne Islands off the Northumberland coast, northern England July 8, 2013. The Farne Islands, which lie off the coast of northeast England, are home to a huge array of wildlife. The islands are owned and protected by the British conservation charity, the National Trust, which says they host some 23 species of seabird, as well as a substantial colony of grey seals, who come to have their pups there in the autumn. Every five years the National Trust carries out a census of the islands' population of puffins, and this year's survey showed there were almost 40,000 nesting pairs on the islands – an 8 percent rise from 2008. (Photo by Nigel Roddis/Reuters)

Respect, Kronotsky nature reserve, Russia. The photographer’s cat, Ryska – her name means little lynx in Russian – stands outside their cabin and with aggressive posturing warns off a fox. In winter, foxes would regularly visit the cabin searching for food. If one peered in at the window, possible when the snow was deep, Ryska would sit on the other side, fur raised, and growl. When outside, she would hold her ground. The foxes were not always frightened and so encounters could be a sort of dance. (Photo by Igor Shpilenok/Unforgettable Behaviour/NHM)

“Eye of a toad”. Animal Portraits, Łukasz Bożycki, Poland. Early spring sees a pond near Łukasz’s home city of Warsaw, Poland, full of mating frogs and a few toads. On this March day, Łukasz shared the pond with them for an evening, sitting in the icy water in his chest-high waders, keeping as still as possible, despite the numbing cold, so that the amphibians could get used to him. “I wanted to find a fresh way of portraying the amphibians”, he says, “at water level”. Using a telephoto lens, he focused on one lone toad and waited for the sun to dip almost below the horizon before pressing the shutter, using flash to bring out the details in the shadow. His prize was “the glorious pool of sunset colour” and fiery glow of the toad’s eye. Nikon D80 + 70-300mm f4.5-5.6 lens + extension tube; 1/125 sec at f9 (-2.3 e/v); ISO 100; built-in flash. (Photo by Łukasz Bożycki)

Tim Laman is a field biologist and wildlife photojournalist. His pioneering research in the rain forest canopy in Borneo led to a PhD from Harvard and his first National Geographic article in 1997. Since then, he has pursued his passion for exploring wild places and documenting little-known and endangered wildlife by becoming a regular contributor to National Geographic. He has eighteen articles to his credit to date, all of which have had a conservation message. Some have focused on endangered species such as Orangutans or Hornbills, while others, such as a series of articles on Conservation International’s Biodiversity Hotspots, have highlighted regions under intense pressure.