Coast and marine winner: Storm Gull (Lesser black-backed gull), New Haven, East Sussex. (Photo by Craig Denford/British Wildlife Photography Awards)

A frog appears to have a big smile for the camera, Russia. Things are heating up at the Comedy Wildlife Photography Awards as the shortlisted final 40 entries are revealed. This year's competition has featured over 2200 hilarious entries from around the world with photos including a fox face planting in the snow, a dancing brown bear and a pair of seemingly headless penguins – all beautifully photographed with perfect comedy timing and a strong conservation message. (Photo by Artyom Krivosheev/Barcroft Images/Comedy Wildlife Photography Awards 2016)

“Romantic”. A brown bear in Martinselkonen, Finland. (Photo by Valtteri Mulkahainen/Comedy Wildlife Photography Awards)

A golden huddle by Minqiang Lu, China. Two females and a male golden snub-nosed monkey huddle together to keep warm in the extreme cold. Threatened by forest loss and fragmentation, this endangered species is confined to central China. Restricted to living high up in the temperate forests, these monkeys – here in the Qinling mountains in Shaanxi province – feed mostly in the trees, on leaves, bark, buds and lichen. In heavy wind and snow, Minqiang walked up the mountain carrying his equipment. He stayed for half an hour in temperatures of –10C opposite the tree where the group was huddled before achieving this eye-level composition. (Photo by Minqiang Lu/Wildlife Photographer of the Year)

Ten finalists capture the theme of “through young eyes” in this young photographers’ competition that aims to engage youth around the world in wildlife conservation. Here: Heavy bodied jumping spider by Md Rashuidul Rabby, age 23, from Bangladesh. (Photo by Md Rashuidul Rabby/WWD2017)

The fourth annual BigPicture Natural World Photography Competition aims to celebrate the diversity of life on Earth, and encourages people to protect and conserve it. Here: “The Salmon Catchers”. Terrestrial Wildlife. To capture this view of a mother grizzly bear and her cub, photographer Peter Mather set up a camera trap on a log that he knew the bears tended to traverse while fishing for salmon, in the Yukon River watershed in Canada. (Photo by Peter Mather/BigPicture Natural World Photography Competition 2017)

“Family disagreement”. (Photo by Vlado Pirsa/Comedy Wildlife Photo Awards 2019)

Hidden Britain category winner. Garden Spider by Alan Smith from Reading, Berkshire. (Photo by Alan Smith/British Wildlife Photography Awards/PA Wire Press Association)