Highly commended birds : Crested guan by Tim Hunt (UK). ‘This photo shows a crested guan in the cloud forests of Costa Rica as it pauses while foraging on a lone branch. Due to the clouds that are so typical for this habitat, I could often only photograph the bird’s silhouette against a grey sky. But then, for a short moment only, the sun broke through the clouds, and I overexposed the image by over two stops in order to blow out the background and allow this beautifully marked bird to stand out’. (Photo by Tim Hunt/GDT European Wildlife Photographer of the Year 2015)

Highly commended, mammals: Gelada after the storm – Marco Gaiotti (Italy). “Gelada baboons are the only monkey species in the world that feed on grasses. They are native to the tableland of Ethiopia. Every morning large family groups wander from their sleeping places in the steep rock face, up to 1,000 metres high, to the feeding grounds at the tablelands. This image clearly depicts their feeding strategy: they pull out bunches of grass, sort the stalks and then lift them to their mouth. This shot was taken towards the end of the rainy season after a heavy storm”. (Photo by Marco Gaiotti/2019 GDT European Wildlife Photographer of the Year)

Behaviour: Mammals category. Giant Gathering by Tony Wu, USA. Dozens of sperm whales mingled noisily off Sri Lanka’s northeast coast, stacked as far down as Tony could see. This was a congregation of dozens of social units, like a gathering of the clans. Aggregations like this could be a critical part of the whales’ rich social lives but are rarely reported. Some two thirds of the population was wiped out before commercial whaling was banned in 1986. This kind of major gathering could be “a sign that populations are recovering”, says Tony. (Photo by Tony Wu/Wildlife Photographer of the Year 2017)

“Romantic”. A brown bear in Martinselkonen, Finland. (Photo by Valtteri Mulkahainen/Comedy Wildlife Photography Awards)

A golden huddle by Minqiang Lu, China. Two females and a male golden snub-nosed monkey huddle together to keep warm in the extreme cold. Threatened by forest loss and fragmentation, this endangered species is confined to central China. Restricted to living high up in the temperate forests, these monkeys – here in the Qinling mountains in Shaanxi province – feed mostly in the trees, on leaves, bark, buds and lichen. In heavy wind and snow, Minqiang walked up the mountain carrying his equipment. He stayed for half an hour in temperatures of –10C opposite the tree where the group was huddled before achieving this eye-level composition. (Photo by Minqiang Lu/Wildlife Photographer of the Year)

The British Wildlife Photography Awards winners have been revealed, with Lee Acaster from Suffolk taking home the top prize for his shot of a Graylag Goose in London. Acaster, who received £5,000, photographed the animal against an ominous London skyline, with The Shard clearly visible in the background. Here: “Urban Tourist (Graylag Goose)”. Urban category and overall winner. (Photo by Lee Acaster/British Wildlife Photography Awards 2014)

Looking for love by Tony Wu, USA. Highly commended, Animal Portraits. “Accentuating his mature appearance with pastel colours, protruding lips and an outstanding pink forehead, this Asian sheepshead wrasse sets out to impress females and see off rivals, which he will head-butt and bite, near Japan’s remote Sado Island. Individuals start out as females, and when they reach a certain age and size – up to a metre (more than 3 feet) long – can transform into males. Long-lived and slow-growing, the species is intrinsically vulnerable to overfishing”. (Photo by Tony Wu/2018 Wildlife Photographer of the Year)