Winner – Animal Portraits: The pose by Mogens Trolle, Denmark. A young male proboscis monkey cocks his head slightly and closes his eyes. Unexpected pale blue eyelids now complement his immaculately groomed auburn hair. He poses for a few seconds as if in meditation. He is a wild visitor to the feeding station at Labuk Bay Proboscis Monkey Sanctuary in Sabah, Borneo – “the most laid-back character”, says Trolle, “quite unlike anything I’ve ever seen on another monkey” – connects us, he hopes, with a fellow primate. (Photo by Mogens Trolle/Wildlife Photographer of the Year 2020)

Bloodthirsty by Thomas P. Peschak, Germany/South Africa — winner, Behaviour: birds. When rations run short on Wolf Island, in the remote northern Galápagos, the sharp-beaked ground finches become vampires. Their sitting targets are Nazca boobies and other large birds. The finches rely on a scant diet of seeds and insects, which regularly dries up, so they drink blood to survive. ‘I’ve seen more than half a dozen finches drinking from a single Nazca booby,’ says Tom. Rather than leave their nests the boobies tolerate the vampires, and the blood loss doesn’t seem to cause permanent harm. (Photo by Thomas P. Peschak/2018 Wildlife Photographer of the Year)

Shooting star by Tony Wu, US/Japan. Winner, underwater. The electrifying reproductive dance of a giant sea star. As the surrounding water filled with sperm and eggs from spawning sea stars, Wu faced several challenges. Stuck in a small, enclosed bay with only a macro lens for photographing small subjects, he backed up to squeeze the undulating sea star into his field of view. The dancing posture of spawning sea stars rising and swaying may help release eggs and sperm, or may help sweep the eggs and sperm into the currents where they fertilise together in the water. Kinko Bay, Japan. (Photo by Tony Wu/Wildlife Photographer of the Year)

Sultan, a famous captive fennec that is displayed tied on a rope in front of a tourist shop, is the main attraction in the souk of Douz, a desert town in Tunisia. By the display of such a charismatic animal, tourists are often lured to buy things or pay for pictures. On inquiry, although Sultan has been caught as a pup in the wild, the owners of the shop reassure the foreigners stating that the animal is ‘domestic’. (Photo by Bruno D’Amicis/Fritz Pölking Prize/GDT EWPY 2015)

Students take photos with a giant King Kong sculptures made out straw in the northern Thai province of Chiang Mai on April 3, 2019. (Photo by Lillian Suwanrumpha/AFP Photo)

A lizard is seen hunting for fish during the daytime on May 04, 2017 in West Sumatra, Indonesia. (Photo by Riau Images/Barcroft Images)

A pitch invader is seen on the pitch during the UEFA Euro 2020 Championship Group B match between Finland and Belgium at Saint Petersburg Stadium on June 21, 2021 in Saint Petersburg, Russia. (Photo by Kirill Kudryavtsev/Reuters)

A “creuseur”, or digger, a plastic lantern on his head, readies to enter a copper and cobalt mine in Kawama, Democratic Republic of Congo on June 8, 2016. Cobalt is used in the batteries for electric cars and mobile phones. Working conditions are dangerous, often with no safety equipment or structural support for the tunnels. The diggers say they are paid on average US$2-3/day. (Photo by Michael Robinson Chavez/The Washington Post)