Shortlisted: Sólheimasandur aircraft crash site, Iceland by Ollie Conway. (Photo by Ollie Conway/Historic Photographer of the Year Awards 2019/The Guardian)

Aurora Australis from Beerbarrel Beach, by James Stone. Runner Up: Aurorae. (Photo by James Stone/Astronomy Photographer of the Year)

Birds behaviour winner: Land of the Eagle by Audun Rikardsen, Norway. High on a ledge, on the coast near his home in northern Norway, Rikardsen carefully positioned an old tree branch that he hoped would make a perfect golden eagle lookout. To this, he bolted a tripod head with a camera, flashes and motion sensor attached, and built himself a hide a short distance away. From time to time, he left road‑kill carrion nearby. Very gradually – over the next three years – a golden eagle got used to the camera and started to use the branch regularly to survey the coast below. (Photo by Audun Rikardsen/2019 Wildlife Photographer of the Year)

British Waters Wide Angle category runner-up. Grass snake swimming along a garden pond by Jack Perks (UK)in Nottinghamshire, UK. “I’m always on the lookout for unusual freshwater subjects and grass snakes are a species I’ve been after for years. I was told about a pond where the odd grass snake hangs around the lilly pads for frogs. I put my drysuit on and got into the water and could see one slithering along the surface. Slowly making my way towards it with my head only just poking above I got the spilt shot”. (Photo by Jack Perks/Underwater Photographer of the Year 2019)

“The Supercell”. This shortlisted image by Dennis Oswald Huge farmland of was taken in south-west Oklahoma, US. (Photo by Dennis Oswald/2019 Weather Photographer of the Year/RMetS)

Run by the Society of German Nature Photographers (Gesellschaft Deutscher Tierfotografen), the prestigious annual contest celebrates the best wildlife photography from members in Germany. Here: Landscapes, second place: Florian Smit, “Rainforest” (Danum valley, Borneo). (Photo by Florian Smit/2019 GDT Nature Photographer of the Year)

Highly commended, mammals: Gelada after the storm – Marco Gaiotti (Italy). “Gelada baboons are the only monkey species in the world that feed on grasses. They are native to the tableland of Ethiopia. Every morning large family groups wander from their sleeping places in the steep rock face, up to 1,000 metres high, to the feeding grounds at the tablelands. This image clearly depicts their feeding strategy: they pull out bunches of grass, sort the stalks and then lift them to their mouth. This shot was taken towards the end of the rainy season after a heavy storm”. (Photo by Marco Gaiotti/2019 GDT European Wildlife Photographer of the Year)

Tropical acrobatics by Adrià López Baucells in Manaus, Brazil. An unidentified South American marsupial, although the characteristic black markings on its face indicate it may be a mouse opossum. These small creatures are nocturnal and feed on bugs, fruit and bird eggs. (Photo by Adrià López Baucells/2019 Royal Society of Biology Photography Competition)