Winner – Animal Portraits: The pose by Mogens Trolle, Denmark. A young male proboscis monkey cocks his head slightly and closes his eyes. Unexpected pale blue eyelids now complement his immaculately groomed auburn hair. He poses for a few seconds as if in meditation. He is a wild visitor to the feeding station at Labuk Bay Proboscis Monkey Sanctuary in Sabah, Borneo – “the most laid-back character”, says Trolle, “quite unlike anything I’ve ever seen on another monkey” – connects us, he hopes, with a fellow primate. (Photo by Mogens Trolle/Wildlife Photographer of the Year 2020)

I had to stay late at work, Chubut, Argentina. “South sea elephant in Patagonia (Isla Escondida) They adopt very curious gestures!”. (Photo by Luis Burgue/Comedy Wildlife Photography Awards 2020)

A zebra walks through the recently flooded Safari Park located at Safari World November 2, 2011 in Bangkok, Thailand. (Photo by Paula Bronstein /Getty Images)

Nosy neighbour by Sam Hobson, UK. Sam knew exactly who to expect when he set his camera on the wall one summer’s evening in a suburban street in Bristol, the UK’s famous fox city. He wanted to capture the inquisitive nature of the urban red fox in a way that would pique the curiosity of its human neighbours about the wildlife around them. (Photo by Sam Hobson/2016 Wildlife Photographer of the Year)

This year’s overall winner and winner of the coast and marine category is George Stoyle with his image “Hitchhikers” of a Lion’s mane jellyfish, photographed at St Kilda, off the Island of Hirta, Scotland. (Photo by George Stoyle/British Wildlife Photography Awards 2016)

Highly commended birds : Crested guan by Tim Hunt (UK). ‘This photo shows a crested guan in the cloud forests of Costa Rica as it pauses while foraging on a lone branch. Due to the clouds that are so typical for this habitat, I could often only photograph the bird’s silhouette against a grey sky. But then, for a short moment only, the sun broke through the clouds, and I overexposed the image by over two stops in order to blow out the background and allow this beautifully marked bird to stand out’. (Photo by Tim Hunt/GDT European Wildlife Photographer of the Year 2015)

Behaviour: Mammals category. Giant Gathering by Tony Wu, USA. Dozens of sperm whales mingled noisily off Sri Lanka’s northeast coast, stacked as far down as Tony could see. This was a congregation of dozens of social units, like a gathering of the clans. Aggregations like this could be a critical part of the whales’ rich social lives but are rarely reported. Some two thirds of the population was wiped out before commercial whaling was banned in 1986. This kind of major gathering could be “a sign that populations are recovering”, says Tony. (Photo by Tony Wu/Wildlife Photographer of the Year 2017)

Highly commended, mammals: Gelada after the storm – Marco Gaiotti (Italy). “Gelada baboons are the only monkey species in the world that feed on grasses. They are native to the tableland of Ethiopia. Every morning large family groups wander from their sleeping places in the steep rock face, up to 1,000 metres high, to the feeding grounds at the tablelands. This image clearly depicts their feeding strategy: they pull out bunches of grass, sort the stalks and then lift them to their mouth. This shot was taken towards the end of the rainy season after a heavy storm”. (Photo by Marco Gaiotti/2019 GDT European Wildlife Photographer of the Year)