Commended. In late May, about a quarter of a million snow geese arrive from North America to nest on Wrangel Island, in northeastern Russia. They form the world's largest breeding colony of snow geese. Photographer Sergey Gorshkov spent two months on the remote island photographing the unfolding dramas. Arctic foxes take advantage of the abundance of eggs, caching surplus eggs for leaner times. But a goose (here the gander) is easily a match for a fox, which must rely on speed and guile to steal eggs. “The battles were fairly equal”, notes Sergey, “and I only saw a fox succeed in grabbing an egg on a couple of occasions, despite many attempts”. Surprisingly, “the geese lacked any sense of community spirit”, he adds, “and never reacted when a fox harassed a neighboring pair nesting close by”. (Photo by Sergey Gorshkov/Veolia Environnement Wildlife Photographer)

This month’s selection of travel imagery mixes quiet moments and superlative views. The overall 2018 prize is a West Greenland trip with Wild Photography Holidays. Here: “This was taken on a safari in Kruger national park, South Africa. The scene was like something straight out of a storybook: a wild baboon, impala and elephant crossing the road together, all lined up neatly in a row”. (Photo by Will Clarke/The Guardian)

In this handout image provided by the City of New York, a Nissan NV200 is seen as a New York City taxi cab in New York City. The Taxi and Limousine Commission along with New York City Mayor Michael Bloomberg announced the Nissan NV200 designed by Nissan North America, Inc., has been chosen as the winner of the Taxi of Tomorrow competition and will become the City's exclusive taxicab for a decade. (Photo by City of New York via Getty Images)

Runner-up. “The heavens opened and the streets around Shibuya, Tokyo, were suddenly drenched in even more colour and movement. I was taking photos (my friend was holding the umbrella) when I noticed this woman at the crossing. I’m pleased with the futuristic feel of the image”. (Photo by Katherine Bridgestock/The Guardian)

Winner. “I took this vertical image in the Quarry Bay district of Hong Kong during the dusk ‘blue hour’, when there was a perfect balance between the ambient light in the sky and the artificial lights of the high-rise residential buildings”. (Photo by Jatinder Heer/The Guardian)

Behaviour winner; Courting Royals: two royal terns in courtship display by Kristian Bell. Another beautiful morning on a beautiful beach on the Gulf Coast of Florida seemed to prompt these two royal terns to commence an intricate courtship dance. The photograph was taken with a Canon 300mm lens and 2x extender. (Photo by Kristian Bell/Deakin University/Royal Society Publishing Photography Competition 2018)

Honourable mention, Behaviour category. Toss the scorpion – Indian roller playing with its kill by Susmita Datta. The image was taken during an early morning safari drive at Tadoba Andhari Tiger Reserve in India. (Photo by Susmita Datta/PA Wire/Royal Society Publishing Photography Competition 2017)