Winner. “I took this vertical image in the Quarry Bay district of Hong Kong during the dusk ‘blue hour’, when there was a perfect balance between the ambient light in the sky and the artificial lights of the high-rise residential buildings”. (Photo by Jatinder Heer/The Guardian)

Behaviour winner; Courting Royals: two royal terns in courtship display by Kristian Bell. Another beautiful morning on a beautiful beach on the Gulf Coast of Florida seemed to prompt these two royal terns to commence an intricate courtship dance. The photograph was taken with a Canon 300mm lens and 2x extender. (Photo by Kristian Bell/Deakin University/Royal Society Publishing Photography Competition 2018)

Runner-up. “The heavens opened and the streets around Shibuya, Tokyo, were suddenly drenched in even more colour and movement. I was taking photos (my friend was holding the umbrella) when I noticed this woman at the crossing. I’m pleased with the futuristic feel of the image”. (Photo by Katherine Bridgestock/The Guardian)

Tropical acrobatics by Adrià López Baucells in Manaus, Brazil. An unidentified South American marsupial, although the characteristic black markings on its face indicate it may be a mouse opossum. These small creatures are nocturnal and feed on bugs, fruit and bird eggs. (Photo by Adrià López Baucells/2019 Royal Society of Biology Photography Competition)

These stunning photographs of the world’s last remaining Mongolian eagle keepers show the incredible bond between man and bird. The images were taken by photography tour guide Daniel Kordan, 29, in September 2018. (Photo by Daniel Kordan/Caters News Agency)

“Scott Linstead is an internationally published, freelance wildlife photographer/writer. His clients include Natural History Magazine, Hewlett Packard, Ranger Rick Magazine and a number of wildlife publications in North America and Europe. Scott's column on the techniques of bird photography appears in every issue of Outdoor Photography Canada”.

Photo: A veiled chameleon extends its tongue to catch a cricket. Canadian wildlife photographer Scott Linstead, formerly an aerospace engineer and high school teacher, uses a device called Phototrap “to not only photograph the elusive, but also the unimaginably quick”. (Photo by Scott Linstead)

Birds are seen at Kizilirmak Bird Paradise in Turkey's Samsun on April 22, 2021. Kizilirmak Delta Bird Sanctuary, which is on the UNESCO World Heritage Tentative List and hosts 359 to 487 species of birds in Turkey, became more active with the arrival of spring. (Photo by Mehmet Kumcagiz/Anadolu Agency via Getty Images)

A bird vendor holds a cage with a bird on a sideroad after a heavy snowfall in Kabul on January 5, 2022. (Photo by Mohd Rasfan/AFP Photo)