Birds behaviour winner: Land of the Eagle by Audun Rikardsen, Norway. High on a ledge, on the coast near his home in northern Norway, Rikardsen carefully positioned an old tree branch that he hoped would make a perfect golden eagle lookout. To this, he bolted a tripod head with a camera, flashes and motion sensor attached, and built himself a hide a short distance away. From time to time, he left road‑kill carrion nearby. Very gradually – over the next three years – a golden eagle got used to the camera and started to use the branch regularly to survey the coast below. (Photo by Audun Rikardsen/2019 Wildlife Photographer of the Year)

A lone male cheetah is set upon by a pack of African wild dogs. Peter Haygarth had been following the dogs as they hunted in Zimanga Private Game Reserve in KwaZulu-Natal, South Africa. On first encountering the cheetah, the dogs were wary, but as the rest of the pack arrived, their confidence grew and they began to encircle the cat. Peter kept his focus on the cat’s face. In a few minutes the spat was over as the cheetah fled. (Behaviour: mammals category). (Photo by Peter Haygarth)

The National Geographic Photo Ark is a travelling exhibition of photographer Joel Sartore’s quest to create a photo archive of biodiversity around the world. So far, Sartore has captured studio portraits of more than 6,000 species – a number that he hopes to double. On 1 July, the ark will open at Melbourne zoo – the first time it has been exhibited in the southern hemisphere. More than 50 portraits will be on display, including many of Australian endangered animals being protected by programs at the zoo itself. These captions have been edited from text supplied by Melbourne zoo. Here: Barking owl. (Photo by Joel Sartore/National Geographic Photo Ark/The Guardian)

A frog appears to have a big smile for the camera, Russia. Things are heating up at the Comedy Wildlife Photography Awards as the shortlisted final 40 entries are revealed. This year's competition has featured over 2200 hilarious entries from around the world with photos including a fox face planting in the snow, a dancing brown bear and a pair of seemingly headless penguins – all beautifully photographed with perfect comedy timing and a strong conservation message. (Photo by Artyom Krivosheev/Barcroft Images/Comedy Wildlife Photography Awards 2016)

Prepare yourself for some rib-tickling laughter because the Comedy Wildlife Awards has announced its finalists. Founded by Tanzania-based photographers Paul Joynson-Hicks MBE and Tom Sullam, the aim of the awards is to put a spotlight on wildlife conservation efforts while simultaneously injecting some humour into the world of wildlife photography. Here: Mountains Gorilla is making grimaces, as he came out of the bush after the rain, in Virunga National Park, Rwanda. (Photo by Josef Friedhuber/Comedy Wildlife Photography Awards/Barcroft Media)

Coast and marine winner: Storm Gull (Lesser black-backed gull), New Haven, East Sussex. (Photo by Craig Denford/British Wildlife Photography Awards)

Arctic treasure by Sergey Gorshkov (Russia). An arctic fox carries its egg trophy from a raid on a snow goose nest and heads for a suitable burial spot. Finalist 2017, Animal Portraits. (Photo by Sergey Gorshkov/2017 Wildlife Photographer of the Year)

A tale of two foxes. Surprising behaviour, witnessed in Wapusk national park, on Hudson Bay, Canada, in early winter. Red foxes don’t actively hunt Arctic foxes, but where the ranges of two predators overlap, there can be conflict. Though the light was poor, the snow-covered tundra provided the backdrop for the moment that the red fox paused with the smaller fox in its mouth in a grim pose. (Photo by Don Gutoski/2015 Wildlife Photographer of the Year)