“Eye of a toad”. Animal Portraits, Łukasz Bożycki, Poland. Early spring sees a pond near Łukasz’s home city of Warsaw, Poland, full of mating frogs and a few toads. On this March day, Łukasz shared the pond with them for an evening, sitting in the icy water in his chest-high waders, keeping as still as possible, despite the numbing cold, so that the amphibians could get used to him. “I wanted to find a fresh way of portraying the amphibians”, he says, “at water level”. Using a telephoto lens, he focused on one lone toad and waited for the sun to dip almost below the horizon before pressing the shutter, using flash to bring out the details in the shadow. His prize was “the glorious pool of sunset colour” and fiery glow of the toad’s eye. Nikon D80 + 70-300mm f4.5-5.6 lens + extension tube; 1/125 sec at f9 (-2.3 e/v); ISO 100; built-in flash. (Photo by Łukasz Bożycki)

Tim Laman is a field biologist and wildlife photojournalist. His pioneering research in the rain forest canopy in Borneo led to a PhD from Harvard and his first National Geographic article in 1997. Since then, he has pursued his passion for exploring wild places and documenting little-known and endangered wildlife by becoming a regular contributor to National Geographic. He has eighteen articles to his credit to date, all of which have had a conservation message. Some have focused on endangered species such as Orangutans or Hornbills, while others, such as a series of articles on Conservation International’s Biodiversity Hotspots, have highlighted regions under intense pressure.

Polar frame, by Dmitry Kokh, Russia. When Kokh’s boat approached the small island of Kolyuchin in the Russian High Arctic, which had been abandoned by humans since 1992, he was surprised to spot movement in one of the houses. Binoculars revealed polar bears – more than 20 in total – exploring the ghost town. Dmitry used a low-noise drone to document them. (Photo by Dmitry Kokh/Wildlife Photographer of the Year 2022)

Arctic treasure by Sergey Gorshkov (Russia). An arctic fox carries its egg trophy from a raid on a snow goose nest and heads for a suitable burial spot. Finalist 2017, Animal Portraits. (Photo by Sergey Gorshkov/2017 Wildlife Photographer of the Year)

A tale of two foxes. Surprising behaviour, witnessed in Wapusk national park, on Hudson Bay, Canada, in early winter. Red foxes don’t actively hunt Arctic foxes, but where the ranges of two predators overlap, there can be conflict. Though the light was poor, the snow-covered tundra provided the backdrop for the moment that the red fox paused with the smaller fox in its mouth in a grim pose. (Photo by Don Gutoski/2015 Wildlife Photographer of the Year)

Three elephant seals put on a show in Roie Galitz's “Three Tanors”, taken on January 7, 2016 in South Georgia Island. The Comedy Wildlife Photography Awards are in full swing, so check out some of the fierce competitors jostling for the top prize this year. Photographers Paul Joynson-Hicks MBE and Tom Sullam founded the awards to spotlight wildlife conservation efforts and to inject some humour into the world of wildlife photography. (Photo by Roie Galitz/CWPA/Barcroft Images)

Andrew Parkinson, animal behaviour category winner: Crepuscular Contentment, Derbyshire. “In 15 years of working with badgers I’ve never seen a badger sit out in the open to have a scratch. I was sat concealed behind a tree and downwind so it was especially nice that the badger had his back to me, demonstrating just how inconspicuous and inconsequential my presence was”. (Photo by Andrew Parkinson/British Wildlife Photography Awards 2017)

A comedic photo of a cheetah appearing to find something hilarious by Dutton Robert for the Comedy Wildlife Photo Awards 2016, South Africa, April, 2012. (Photo by Dutton Robert/Barcroft Images/Comedy Wildlife Photo Awards)