A tale of two foxes. Surprising behaviour, witnessed in Wapusk national park, on Hudson Bay, Canada, in early winter. Red foxes don’t actively hunt Arctic foxes, but where the ranges of two predators overlap, there can be conflict. Though the light was poor, the snow-covered tundra provided the backdrop for the moment that the red fox paused with the smaller fox in its mouth in a grim pose. (Photo by Don Gutoski/2015 Wildlife Photographer of the Year)

Touching and dramatic portraits and landscape shots have won prizes at Australia's prestigious photography prize. Photo: Winner of the NSW (New South Wales) prize: Peter Solness said: “I wanted to re-imagine the lost waterways, so I got my light-painting tools to work. In this image, water is being released from the top of the historic Centennial Park No. 2 Reservoir, which was built in 1925 and holds 90 megalitres of water. After 89 years of incarceration these waters now run free!”. (Photo by Peter Solness/Head On)

Wakodahatchee wetlands, Delray Beach, Florida, US. Equipped with sinewy necks and spear-like bills, great blue herons can lunge with fearsome speed to strike their aquatic prey. Adults will also employ rapid stabbing motions as one aspect of their complex courtship displays; they’re seemingly dangerous moves, but fitting to the intensity of mating season. (Photo by Melissa Rowell/Audubon Photography Awards)

Portrait category, bronze award winner. Purple-crested Turaco Gallirex porphyreolophus. Lower Mpushini, near Pietermaritzburg, KwaZulu-Natal, South Africa. (Photo by Richard Flack/Bird Photographer of the Year 2022)

Brain-on-a-chip. Dazzling in green and magenta this image shows the nerve fibres (in green) produced by neural stem cells (in magenta) as they grow on a synthetic gel. Captured by a technique known as confocal microscopy, the image is part of research shedding light on how tinkering with the environment can affect the way in which nerve fibres grow. (Photo by Collin Edington and Iris Lee/Massachusetts Institute of Technology/Wellcome Images)

Birds behaviour winner: Land of the Eagle by Audun Rikardsen, Norway. High on a ledge, on the coast near his home in northern Norway, Rikardsen carefully positioned an old tree branch that he hoped would make a perfect golden eagle lookout. To this, he bolted a tripod head with a camera, flashes and motion sensor attached, and built himself a hide a short distance away. From time to time, he left road‑kill carrion nearby. Very gradually – over the next three years – a golden eagle got used to the camera and started to use the branch regularly to survey the coast below. (Photo by Audun Rikardsen/2019 Wildlife Photographer of the Year)

“The power of nature”. Magma, ash and gas erupt from Mount Etna in December 2015, rising to a height of several kilometres. Winner: Nature. (Photo by Giuseppe Mario Famiani/SIPA Contest)

A cave diver in Anhumas Abyss, Bonito, Brazil. (Photo by Marcio Cabral/Epson International Pano Awards 2018)